Forests are home to some of the most complex microbial communities (Fierer et al. 2012) which drive biogeochemical cycles (Clemmensen et al. 2013; van der Heijden et al. 2008) and account for substantial terrestrial biomass (Nielsen et al. 2011). Fungi, through their ecological roles as decomposers, mutualists, or pathogens, are particularly important in breaking down organic matter and mediating plant nutrition. Despite this, little is known about the variability, composition, and structure of forest soil fungal communities (Tedersoo et al. 2014). In fact, much of global fungal diversity remains undocumented (Blackwell 2011; Hawksworth 2012).