US Forest Service Research & Development
Contact Information
  • US Forest Service Research & Development
  • 1400 Independence Ave., SW
  • Washington, D.C. 20250-0003
  • 800-832-1355
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Priority Areas

  • Forest Disturbances

SRS Science Area

Experimental Forests & Ranges

John Willis

Research Forester
521 Devall Drive
United States

Phone: 334-826-8700 x133
Fax: 334-821-0037
Contact John Willis

Current Research

  • Longleaf pine restoration on xeric sites
  • Accelerating late-successional forest structure in the Pacific Northwest
  • Retaining forest flammability in upland hardwood stands
  • Mixed-stand dynamics in the southeastern United States
  • Gap dynamics in northern hardwoods        

Research Interests

  • Restoring degraded ecosystems
  • Understanding contemporary stand dynamics
  • Developing silvicultural strategies for maintaining species diversity


  • Michigan State University, Ph.D. Forestry and Evolutionary Ecology and Behavior Biology Forest Ecology and Silviculture 2014
  • Colorado State University, M.S. Forest Science Silviculture 2009
  • Colorado State University, B.S Forestry Fire Science 2007

Awards & Recognition

  • Manierre Award, 2019
    Award for research conducted with the Huron Mountain Wildlife Foundation
  • Silviculture Prize for Best Publication (co-author), 2018
    Institute of Chartered Foresters


Citations of non US Forest Service Publications

  • -Willis, J.L., Schnake, D.K., Wetzstein, B., Yow, J., Guinto, D., Ulrich, S., DePerno, C.S., Lashley, M.A. (2019). Seed depredation negates the benefits of midstory hardwood removal on longleaf pine seedling establishment. Restoration Ecology. 0, 1-9


    -Willis, J.L., Walters, M.B. (2018). Nutrition and mycorrhizae affect interspecific patterns of seedling growth on coarse wood and mineral soil substrates. Ecosphere. 9, 1-17


    -Webster, C.R., Dickinson, Y.L. Burton, J.I, Frelich, L.E., Jenkins, M.A., Kern, C.C., Raymond, P., Saunders, M.R., Walters, M.B., Willis, J.L. (2018). Promoting and maintaining diversity in contemporary hardwood forests: confronting contemporary drivers of change and the loss of ecological memory. Forest Ecology and Management. 421, 98-108


    -Willis, J.L., Roberts, S.D., Harrington, C.A. (2018). Variable density thinning promotes variable structural responses 14 years after treatment in the Pacific Northwest. Forest Ecology and Management. 410, 114-125


    -Kern, C.C., Burton, J.I., Raymond, P., D’Amato, A.W., Keeton, W.S., Royo, A.A., Walters, M.B., Willis, J.L. (2017). Challenges facing gap-based silviculture and possible solutions for mesic northern forests in North America. Forestry. 89, 1-14


    -Walters, M.B., Farinosi, E., Willis, J.L. (2016). Managing for diversity: harvest gap size drives complex light, vegetation, and deer herbivory impacts on tree seedlings. Ecosphere. 7, 1-29

    -Willis, J.L, Walters, M.B. (2016). Local seed source availability limits young seedling populations for some species more than other factors in northern hardwood forests. Forest Science. 62, 440-448


    -Willis, J.L., Walters, M.B., Gottschalk, K.W. (2015). Scarification and harvest gap size have interacting effects on young seedling establishment. Forest Ecology and Management. 347, 237-246


    -Walters, M.B., Willis, J. L, Gottschalk, K.W. (2014). Can light and nitrate vs. ammonium growth responses among mixed hardwood-conifer forest species be predicted by their ecologies and help explain community scale tree diversity? Canadian Journal of Forest Research. 44, 1356-1368

Last updated on : 10/12/2021