Current Research

Soil ecology, biology and biogeochemistry

Research Interests

Soils, decay, nutrient cycling, soil organisms

Past Research

Earthworm ecology, soil biology and biogeochemistry

Education

• Department of Environmental, Population and Organismic (EPO) Biology, University of Colorado-Boulder, Ph.D. Soil ecology and biology. Dissertation: Soil fauna, microbes and plant litter decomposition in tropical and subalpine forests. 1999
• Department of Biology, University of Puerto Rico, M.Sc. Soil ecology. Thesis: Earthworm abundance and distribution pattern in two plant communities within a subtropical wet forest of Puerto Rico. 1996
• Department of Biology, University of Puerto Rico, B.S. Biology 1993

Professional Experience

Professional Organizations

• National Inventory Monitoring And Assessment Steering Committee Of Usfs Nfs, Member (2014 - Current)
  Washington DC
• Special Working Group, Conservation Areas In Pr, Member (2014 - Current)
  Special Working Group for Commonwealth Government of Puerto Rico for Strategic Augmentation of Conservation Areas in Puerto Rico
• Luquillo Critical Zone Observatory, Co-Principal Investigator (2013 - Current)
  US NSF Award to Univ. of New Hampshire, EAR 1331841 ($5M) Luquillo CZO: The role of hot spots and hot moments in tropical landscape evolution and functioning of the critical zone (Sept 2013 to Aug 2018)
• El Yunque National Forest Management Plan, Interdisciplinary Team, Science Advisor And Member (2012 - Current)
  Interdisciplinary Team of El Yunque National Forest Management Plan
• Luquillo Long Term Ecological Research, Co-Principal Investigator (2010 - Current)
  US NSF Award to Univ. of Puerto Rico, DEB 1239764 ($1.96M), Understanding Change in the Ecosystems of Northeastern Puerto Rico (Jan 2014 to Dec 2015)
• Albuquerque Service Center Operations Customer Service Board, Usfs National R&D Project Leader Representative (2012 - 2016)
• International Scholarly Research Network-Soil Science, Editorial Board Member (2011 - 2011)

Awards & Recognition

Featured Publications & Products

• Dalling, James W.; Heineman, Katherine; Gonzalez, Grizelle; Ostertag, Rebecca. 2016. Geographic, environmental and biotic sources of variation in the nutrient relations of tropical montane forests.
• Shiels, Aaron B.; Gonzalez, Grizelle; Lodge, D. Jean; Willig, Michael R; Zimmerman, Jess K. 2015. Cascading Effects of Canopy Opening and Debris Deposition from a Large-Scale Hurricane Experiment in a Tropical Rain Forest.
• Gonzalez, G.; Lodge, D.J.; Richardson, B.A.; Richardson, M.J. 2014. A canopy trimming experiment in Puerto Rico: The response of litter decomposition and nutrient release to canopy opening and debris deposition in a subtropical wet forest.
• Gonzalez, G.; Waide, R.B.; Willig, M.R. 2013. Advancements in the understanding of spatiotemporal gradients in tropical landscapes: a Luquillo focus and global perspective.
• Gonzalez, Grizelle; Huang, C.Y.; Chang, S.C. 2008. Earthworms and post agricultural succession in the Neotropics.
• Gonzalez, Grizelle; Yu Huang, Ching; Zou, Xiaoming; Rodriguez, Carlos. 2006. Earthworm invasions in the tropics.

Publications

• Miller, Paul W.; Mote, Thomas L.; Ramseyer, Craig A.; Van Beusekom, Ashley; Scholl, Martha; Gonzalez, Grizelle. 2018. A 42 year inference of cloud base height trends in the Luquillo Mountains of northeastern Puerto Rico.
• Djukic, Ika; Kepfer-Rojas, Sebastian; Schmidt, Inger Kappel; Larsen, Klaus Steenberg; Beier, Claus; Berg, Björn; Verheyen, Kris; Caliman, Adriano; Paquette, Alain; Gutiérrez-Girón, Alba; Humber, Alberto; Valdecantos, Alejandro; Petraglia, Alessandro; Alexander, Heather; Augustaitis, Algirdas; Saillard, Amélie; Fernández, Ana Carolina Ruiz; Sousa, Ana I.; Lillebø, Ana I.; da Rocha Gripp, Anderson; Francez, André-Jean; Fischer, Andrea; Bohner, Andreas; Malyshev, Andrey; Andrić, Andrijana; Smith, Andy; Stanisci, Angela; Seres, Anikó; Schmidt, Anja; Avila, Anna; Probst, Anne; Ouin, Annie; Khuroo, Anzar A.; Verstraeten, Arne; Palabral-Aguilera, Arely N.; Stefanski, Artur; Gaxiola, Aurora; Muys, Bart; Bosman, Bernard; Ahrends, Bernd; Parker, Bill; Sattler, Birgit; Yang, Bo; Juráni, Bohdan; Erschbamer, Brigitta; Ortiz, Carmen Eugenia Rodriguez; Christiansen, Casper T.; Carol Adair, E.; Meredieu, Céline; Mony, Cendrine; Nock, Charles A.; Chen, Chi-Ling; Wang, Chiao-Ping; Baum, Christel; Rixen, Christian; Delire, Christine; Piscart, Christophe; Andrews, Christopher; Rebmann, Corinna; Branquinho, Cristina; Polyanskaya, Dana; Delgado, David Fuentes; Wundram, Dirk; Radeideh, Diyaa; Ordóñez-Regil, Eduardo; Crawford, Edward; Preda, Elena; Tropina, Elena; Groner, Elli; Lucot, Eric; Hornung, Erzsébet; Gacia, Esperança; Lévesque, Esther; Benedito, Evanilde; Davydov, Evgeny A.; Ampoorter, Evy; Bolzan, Fabio Padilha; Varela, Felipe; Kristöfel, Ferdinand; Maestre, Fernando T.; Maunoury-Danger, Florence; Hofhansl, Florian; Kitz, Florian; Sutter, Flurin; Cuesta, Francisco; de Almeida Lobo, Francisco; de Souza, Franco Leandro; Berninger, Frank; Zehetner, Franz; Wohlfahrt, Georg; Vourlitis, George; Carreño-Rocabado, Geovana; Arena, Gina; Pinha, Gisele Daiane; González, Grizelle; Additional, Authors; Yesilonis, Ian; Additional, Authors ; Rustad, Lindsey; et, al. 2018. Early stage litter decomposition across biomes.
• Quiñones, Maya; Parés-Ramos, Isabel K.; Gould, William A.; Gonzalez, Grizelle; McGinley, Kathleen; Ríos, Pedro. 2018. El Atlas del Bosque Nacional El Yunque.
• Quiñones, Maya; Parés-Ramos, Isabel K.; Gould, William A.; Gonzalez, Grizelle; McGinley, Kathleen; Ríos, Pedro. 2018. El Yunque National Forest Atlas.
• Van Beusekom, Ashley E.; Gould, William A.; Monmany, A. Carolina; Khalyani, Azad Henareh; Quiñones, Maya; Fain, Stephen J.; Andrade-Núñez, Maria José; González, Grizelle. 2018. Fire weather and likelihood: characterizing climate space for fire occurrence and extent in Puerto Rico.
• Franklin, Janet; Andrade, Riley; Daniels, Mark L.; Fairbairn, Patrick; Fandino, Maria C.; Gillespie, Thomas W.; González, Grizelle; Gonzalez, Otto; Imbert, Daniel; Kapos, Valerie; Kelly, Daniel L.; Marcano-Vega, Humfredo; Meléndez-Ackerman, Elvia J.; McLaren, Kurt P.; McDonald, Morag A.; Ripplinger, Julie; Rojas-Sandoval, Julissa; Ross, Michael S.; Ruiz, Jorge; Steadman, David W.; Tanner, Edmund V. J.; Terrill, Inge; Vennetier, Michel. 2018. Geographical ecology of dry forest tree communities in the West Indies.
• Van Beusekom, Ashley ; Alvarez-Berríos, Nora ; Gould, William ; Quiñones, Maya ; González, Grizelle . 2018. Hurricane Maria in the U.S. Caribbean: Disturbance Forces, Variation of Effects, and Implications for Future Storms.
• Feng, Xiaohui; Uriarte, María; González, Grizelle; Reed, Sasha; Thompson, Jill; Zimmerman, Jess K.; Murphy, Lora. 2018. Improving predictions of tropical forest response to climate change through integration of field studies and ecosystem modeling.
• Kimball, Bruce A.; Alonso-Rodríguez, Aura M.; Cavaleri, Molly A.; Reed, Sasha C.; González, Grizelle; Wood, Tana E. 2018. Infrared heater system for warming tropical forest understory plants and soils.
• Liu, Xianbin ; Zeng, Xiucheng ; Zou, Xiaoming ; González, Grizelle ; Wang, Chao ; Yang, Si . 2018. Litterfall Production Prior to and during Hurricanes Irma and Maria in Four Puerto Rican Forests.
• Liu, Xianbin ; Zeng, Xiucheng ; Zou, Xiaoming ; Lodge, Deborah ; Stankavich, Sarah ; González, Grizelle ; Cantrell, Sharon . 2018. Responses of Soil Labile Organic Carbon to a Simulated Hurricane Disturbance in a Tropical Wet Forest.
• Richardson, Barbara; Richardson, Michael; González, Grizelle. 2018. Responses of Two Litter-Based Invertebrate Communities to Changes in Canopy Cover in a Forest Subject to Hurricanes.
• Prather, Chelse M.; Belovsky, Gary E.; Cantrell, Sharon A.; González, Grizelle. 2018. Tropical herbivorous phasmids, but not litter snails, alter decomposition rates by modifying litter bacteria.
• Van Beusekom, Ashley E.; Gonzalez, Grizelle; Scholl, Martha A. 2017. Analyzing cloud base at local and regional scales to understand tropical montane cloud forest vulnerability to climate change.
• Monmany, Ana Carolina; Gould, William; Andrade-Nunez, Maria Jose; Gonzalez, Grizelle; Quinones, Maya. 2017. Characterizing Predictability of Fire Occurrence in Tropical Forests and Grasslands: The Case of Puerto Rico.
• Chen, Dingfang; Yu, Mei; González, Grizelle; Zou, Xiaoming; Gao, Qiong. 2017. Climate impacts on soil carbon processes along an elevation gradient in the tropical Luquillo Experimental Forest.
• Gonzalez, Grizelle. 2017. Deadwood, soil biota and nutrient dynamics in tropical forests: a review of case studies from Puerto Rico.
• Gonzalez, Grizelle; Barberena, María F. 2017. Ecology of soil arthropod fauna in tropical forests: A review of studies from Puerto Rico.
• Rhea-Fournier, Dylan; Gonzalez, Grizelle. 2017. Methodological Considerations in the Study of Earthworms in Forest Ecosystems.
• Murphy, Sheila F.; Stallard, Robert F.; Scholl, Martha A.; Gonzalez, Grizelle; Torres-Sánchez, Angel J. 2017. Reassessing rainfall in the Luquillo Mountains, Puerto Rico: Local and global ecohydrological implications.
• Van Beusekom, Ashley E.; Gonzalez, Grizelle; Rivera, Maria M. 2017. Reply to Comments on Short-Term Precipitation and Temperature Trends along an Elevation Gradient in Northeastern Puerto Rico.
• Lugo, Ariel; Gonzalez, Grizelle. 2017. Sandra Brown (1944 2017): A Distinguished Tropical Ecologist.
• González, Grizelle; Lodge, D. 2017. Soil biology research across latitude, elevation and disturbance gradients: A review of forest studies from Puerto Rico during the past 25 years.
• Melendez-Ackerman, Elvia M.; Rojas-Sandoval, Julissa; Fernandez, Danny S. ; Gonzalez, Grizelle; Lopez, Hana; Sustache, Jose; Morales, Mariely; Garcia-Bermudez, Miguel; Aragon, Susan. 2016. Associations between soil variables and vegetation structure and composition of Caribbean dry forests.
• Gonzalez, Grizelle; Heartsill Scalley, Tamara. 2016. Building a collaborative network to understand regional forest dynamics and advance forestry initiatives in the Caribbean.
• Lodge, D. Jean; Winter, Dirk; Gonzalez, Grizelle; Clum, Naomi. 2016. Effects of hurricane-felled tree trunks on soil carbon, nitrogen, microbial biomass, and root length in a wet tropical forest.
• Heartsill Scalley, Tamara; Gonzalez, Grizelle. 2016. Introduction: Caribbean forest dynamics and regional forestry initiatives.
• Zalamea, Marcela; Gonzalez, Grizelle; Lodge, D. Jean. 2016. Physical, Chemical, and Biological Properties of Soil under Decaying Wood in a Tropical Wet Forest in Puerto Rico.
• Huang, Ching-Yu; Gonzalez, Grizelle; Hendrix, Paul F. 2016. Resource Utilization by Native and Invasive Earthworms and Their Effects on Soil Carbon and Nitrogen Dynamics in Puerto Rican Soils.
• Mercado-Díaz, Joel A.; Gould, William A.; Gonzalez, Grizelle; Lücking, Robert. 2015. Lichens in Puerto Rico: an ecosystem approach.
• Van Beusekom, Ashley; Gonzalez, Grizelle; Rivera, Maria. 2015. Short-term precipitation and temperature trends along an elevation gradient in northeastern Puerto Rico.
• Scholl, Martha A.; Shanley, James B.; Murphy, Sheila F.; Willenbring, Jane K.; Gonzalez, Grizelle. 2015. Stable-isotope and solute-chemistry approaches to flow characterization in a forested tropical watershed, Luquillo Mountains, Puerto Rico.
• Shiels, Aaron; Gonzalez, Grizelle. 2015. Tropical Forest Responses to Large-Scale Experiments.
• Gonzalez, Grizelle. 2015. Puerto Rico: gateway to landscape from an ecological perspective.
• Jennings, Lisa Nicole; Douglas, Jamison; Treasure, Emrys; González, Grizelle. 2014. Climate change effects in El Yunque National Forest, Puerto Rico, and the Caribbean region.
• Silver, W.L.; Hall, S.J.; Gonzalez, Grizelle. 2014. Differential effects of canopy trimming and litter deposition on litterfall and nutrient dynamics in a wet subtropical forest.
• Gonzalez, Grizelle. 2014. Ecologia de las lombrices de tierra.
• Lodge, D. Jean; Cantrell, Sharon A.; Gonzalez, Grizelle. 2014. Effects of canopy opening and debris deposition on fungal connectivity, phosphorus movement between litter cohorts and mass loss.
• Gonzalez, G.; Rivera-Figueroa, F.J.; Gould, W.; Cantrell, S.A.; Pérez-Jiménez, J.R. 2014. Microorganisms in small patterned ground features and adjacent vegetated soils along topographic and climatic gradients in the High Arctic, Canada.
• Shiels, A.B.; Gonzalez, Grizelle; Willig, M.R. 2014. Responses to canopy loss and debris deposition in a tropical forest ecosystem: Synthesis from an experimental manipulation simulating effects of hurricane disturbance.
• Mercado-Diaz, Joel; Gould, William; Gonzalez, Grizelle. 2014. Soil nutrients, landscape age, and Sphagno-Eriophoretum vaginati plant communities in Arctic moist-acidic Tundra landscapes.
• Shiels, A.B.; Gonzalez, Grizelle. 2014. Understanding the key mechanisms of tropical forest responses to canopy loss and biomass deposition from experimental hurricane effects.
• Gonzalez, G.; Willig, M.R.; Waide, R.B. 2013. Ecological gradient analyses in a tropical landscape: multiple perspectives and emerging themes.
• Mercado-Diaz, J.A.; Gould, W.A.; Gonzalez, G.; Lucking, R. 2013. Four new species of Coenogonium (Ascomycota: Ostropales) from vulnerable forest ecosystems in Puerto Rico.
• Ping, C.-L.; Michaelson, G. J.; Stiles, C. A.; Gonzalez, G. 2013. Soil characteristics, carbon stores, and nutrient distribution in eight forest types along an elevational gradient, eastern Puerto Rico.
• Medina, E.; Gonzalez, G.; Rivera, M. M. 2013. Spatial and temporal heterogeneity of rainfall inorganic ion composition in northeastern Puerto Rico.
• Gonzalez, G.; Luce, M.M. 2013. Woody debris characterization along an elevation gradient in northeastern Puerto Rico.
• Zalamea, Marcela; Gonzalez, Grizelle; Gould, William A. 2012. Comparing litterfall and standing vegetation: assessing the footprint of litterfall traps.
• Gonzalez, Grizelle; Murphy, Christina M.; Belén, Juliana. 2012. Direct and indirect effects of millipedes on the decay of litter of varying lignin content.
• McDowell, W.H.; Scatena, F.N.; Waide, R.B.; Brokaw, N.; Camilo, G.R.; Covich, A.P.; Crowl, T.A.; Gonzalez, G.; Greathouse, E.A.; Klawinski, P.; Lodge, D.J.; Lugo, A.E.; Pringle, C.M.; Richardson, B.A.; Richardson, M.J.; Schaefer, D.A.; Silver, W.L.; Thompson, J.; Vogt, D.J.; Vogt, K.A.; Willig, M.R.; Woolbright, L.L.; Zou, X.; Zimmerman, J.K. 2012. Geographic and ecological setting of the Luquillo Mountains.
• Lugo, A.E.; Scatena, F.N.; Waide, R.B.; Greathouse, E.A.; Pringle, C.M.; Willig, M.R.; Vogt, K.A.; Walker, L.R.; Gonzalez, G.; McDowell, W.H.; Thompson, J.; NO-VALUE, . 2012. Management implications and applications of long-term ecological research.
• Barberena-Arias, María Fernanda; Gonzalez, Grizelle; Cuevas, Elvira. 2012. Quantifying variation of soil arthropods using different sampling protocols: is diversity affected.
• Crowl, T.A.; Brokaw, N.; Waide, R.B.; Gonzalez, G.; Beard, K.H.; Greathouse, E.A.; Lugo, A.E.; Covich, A.P.; Lodge, D.J.; Pringle, C.M.; Thompson, J.; Belovsky, G.E. 2012. When and where biota matter, linking disturbance regime, species characteristics, and dynamics of communities and ecosystems.
• Wall, D.A.; Gonzalez, Grizelle; Simmons, B.L. 2011. Seasonally dry forest soil biodiversity and functioning.
• Richardson, Barbara A.; Richardson, Michael J.; Gonzalez, Grizelle; Shiels, Aaron B.; Srivastava, Diane S. 2010. A canopy trimming experiment in Puerto Rico: the response of litter invertebrate communities to canopy loss and debris deposition in a tropical forest subject to hurricanes.
• Lugo, Ariel E.; Gonzalez, Grizelle. 2010. Changing conditions and changing ecosystems: a long-term regional and transcontinental research approach on invasive species.
• Gould, William; Gonzalez, Grizelle; Walker, D.A.; Ping, C.L. 2010. Commentary. Integrating research, education, and traditional knowledge in ecology: A case study of biocomplexity in the Arctic.
• Lilleskov, Erik; Callaham, Jr. Mac A.; Pouyat, Richard; Smith, Jane E.; Castellano, Michael; Gonzalez, Grizelle; Lodge, D. Jean; Arango, Rachel; Green, Frederick. 2010. Invasive soil organisms and their effects on belowground processes.
• Jain, Theresa B.; Gould, William A.; Graham, Russell T.; Pilliod, David S.; Lentile, Leigh B.; Gonzalez, Grizelle. 2008. A soil burn severity index for understanding soil-fire relations in tropical forests.
• Jain, Theresa B.; Gould, William A.; Graham, Russell T.; Pilliod, David S.; Lentile, Leigh B.; Gonzalez, Grizelle. 2008. A soil burn severity index for understanding soil-fire relations in tropical forests Chinese version.
• D.A., Walker; V.E., Romanovsky; C.L., Ping; G.J., Michaelson; R.P., Daanen; Y., Shur; R.A., Peterson; W.B., Krantz; M.K., Raynolds; Gould, William; Gonzalez, Grizelle; D.J., Nicolsky; C.M., Vonlanthen; A.N., Kade; P., Kuss; A.M., Kelley; C.A., Munger; C.T., Tarnocai; N.V., Matveyeva; F.J.A., Daniels. 2008. Arctic patterned-ground ecosystems: A synthesis of field studies and models along a North American Arctic Transect.
• Meddens, Arjan J. H.; Hudak, Andrew T.; Evans, Jeffrey S.; Gould, William A.; Gonzalez, Grizelle. 2008. Characterizing forest fragments in boreal, temperate, and tropical ecosystems.
• Gonzalez, Grizelle; Gould, William; Hudak, Andrew T.; Hollingsworth, Teresa Nettleton. 2008. Decay of aspen (Populus tremuloides Michx.) wood in moist and dry boreal, temperate, and tropical forest fragments.
• Gould, William A.; Gonzalez, Grizelle; Hudak, Andrew T.; Hollingsworth, Teresa Nettleton; Hollingsworth, Jamie. 2008. Forest structure and downed woody debris in boreal temperate, and tropical forest fragments.
• Zalamea, M.; Gonzalez, G. 2008. Leaf Fall Phenology in a Subtropical Wet Forest in Puerto Rico: from Species to Community Patterns.
• Murphy, Christina M.; Gonzalez, Grizelle; Belén, Juliana. 2008. Ordinal abundance and richness of millipedes (Arthropoda: Diplopoda) in a subtropical wet forest in Puerto Rico.
• Li, Yiqing; Gonzalez, Grizelle. 2008. Soil Fungi and Macrofauna in the Neotropics.
• Rodriguez, Carlos; Borges, S.; Martinez, M.A.; Fragoso, C.; James, S.; Gonzalez, Grizelle. 2007. Biodiversidad y ecolog a de las lombrices de tierra en las islas caribe as.
• Gonzalez, Grizelle; Garcia, Emerita; Cruz, Veronica; Borges, Sonia; Zalamea, Marcela; Rivera, Maria M. 2007. Earthworm communities along an elevation gradient in Northeastern Puerto Rico..
• Gonzalez, Grizelle; Li, Y.; Zou, X. 2007. Effects of post-hurricane fertilization and debris removal on earthworm abundance and biomass in subtropical forests in Puerto Rico.
• Zalamea, Marcela; Gonzalez, Grizelle; Ping, Chien-Lu; Michaelson, Gary. 2007. Soil organic matter dynamics under decaying wood in a subtropical wet forest: effect of tree species and decay stage..
• Zalamea, Marcela; Gonzalez, Grizelle. 2007. Substrate-induced respiration in Puerto Rican soils: minimum glucose amendment.
• Gonzalez, Grizelle; Espinoza, Elianid; Liu, Zhigang; Zou, Xiaoming. 2006. A Fluorescent Marking and Re-count Technique Using the Invasive Earthworm, Pontoscolex corethrurus (Annelida: Oligochaeta).
• Gonzalez, Grizelle. 2006. Earthworms as Invasive Species in Latin America the 2nd Latin American Meeting on Oligochaeta (Earthworm) Ecology and Taxonomy.
• Gonzalez, Grizelle. 2006. Editorial: Earthworms as invasive species in Latin America - the 2nd Latin American Meeting on Oligochaeta (earthworm) ecology and taxonomy.
• Hendrix, P.F.; Baker, G.H.; Callaham, M.A. Jr.; Damoff, G.A.; Fragoso, C.; Gonzalez, G.; James, S.W.; Lachnicht, S.L.; Winsome, T.; Zou, X. 2006. Invasion of exotic earthworms into ecosystems inhabited by native earthworms.
• Hendrix, P. F.; Baker, G. H.; Callaham Jr, M. A.; Damoff, G. A.; Fragoso C., ; Gonzalez, G.; James, S. W.; Lachnicht, S. L.; Winsome, T.; Zou, X. 2006. Invasion of exotic earthworms into ecosystems inhabited by native earthworms.
• Callaham, Mac A., Jr..; Gonzalez, Grizelle; Hale, Cynthia M; Henegham, Liam; Lachnicht, Sharon L.; Zou, Xiaoming. 2006. Policy and management responses to earthworm invasions in North America.
• Gould, W.A.; Gonzalez, G.; Carrero Rivera, G. 2006. Structure and composition of vegetation along an elevational gradient in Puerto Rico..
• Huang, Ching-Yu; Gonzalez, Grizelle; Hendrix, Paul F. 2006. The Re-colonization Ability of a Native Earthworm, Estherella spp., in Forests and Pastures in Puerto Rico.
• Torres, Juan A.; Gonzalez, Grizelle. 2005. Wood Decomposition of Cyrilla racemiflora (Cyrillaceae) in Puerto Rican Dry and Wet Forests: A 13-year Case Study..
• Gonzalez, Grizelle; Seastedt, Timothy R.; Donato, Zugeily. 2003. Earthworms, arthropods and plant litter decomposition in aspen (Populus tremuloides) and lodgepole pine(Pinus contorta) forests in Colorado, USA.
• Gonzalez, Grizelle. 2002. Soil organisms and litter decomposition.
• Zou, X; Gonzalez, Grizelle. 2001. Earthworms in tropical tree plantations: effects of management and relations with soil carbon and nutrient use efficiency.
• Gonzalez, Grizelle; Ley, Ruth E.; Schmidt, Steven K.; Zou, Xiaoming; Seastedt, Timothy R. 2001. Soil ecological interactions: comparisons between tropical and subalpine forests.
• Gonzalez, G.; Seastedt, T.R. 2001. Soil fauna and plant litter decomposition in tropical and subalpine forests.
• Gonzalez, G.; Seastedt, T.R. 2000. Comparison of the abundance and composition of litter fauna in tropical and subalpine forests.
• Gonzalez, G.; Zou, X.; Sabat, A.; Fetcher, N. 1999. Earthworm abundance and distribution pattern in contrasting plant communities within a tropical wet forest in Puerto Rico.
• Gonzalez, G.; Zou, X. 1999. Earthworm influence on N availability and the growth of Cecropia schreberiana in tropical pasture and forest soils.
• Gonzalez, G.; Zou, X. 1999. Plant and litter influences on earthworm abundance and community structures in a tropical wet forest.
• Zou, Xiaoming; Gonzalez, Grizelle. 1997. CHANGES IN EARTHWORM DENSITY AND COMMUNITY STRUCTURE DURING SECONDARY SUCCESSION IN ABANDONED TROPICAL PASTURES.
• Gonzalez, G.; Zou, X.; Borges, S. 1996. Earthworm abundance and species composition in abandoned tropical croplands: comparisons of tree plantations and secondary forests..

Research Highlights

Highlight	Title	Year
IITF-2010-010	A Canopy Trimming Experiment in Puerto Rico: the response of litter invertebrate communities to canopy loss and debris deposition in a tropical forest subject to hurricanes Insects and other invertebrates in leaf litter are important components in decomposition, nutrient cycling, and food webs. As part of larger stu ...	2010
IITF-2012-01	Assessing Litterfall Trap Bias Are traps preferentially representing certain kinds of leaf litter	2012
IITF-2013-026	Ecological Gradient Analyses in Tropical Ecosystems Forest Service scientists recently published a book that contains a comprehensive analysis of ecological gradients in the Luquillo Mountains of ...	2013
IITF-2016-19	Geographic, Environmental, and Biotic Sources of Variation in the Nutrient Relations of Tropical Montane Forests Incorporating other geographic, environmental and biotic variables in tropical montane forests biogeochemistry, might give scientists a more acc ...	2016
IITF-2011-03	Integrating Research, Education, and Traditional Knowledge in Ecology Integrating research and education is a fundamental goal of research institutions and agencies but specific approaches not widely published in e ...	2011
IITF-2016-18	Physical, Chemical, and Biological Properties of Soil Under Decaying Wood in a Tropical Wet Forest in Puerto Rico Decaying wood is related to nutrient cycling through its role as either a sink or source of nutrients; however, at micro scales, what is the eff ...	2016
IITF-2015-84	Post-hurricane Canopy Openings Influence Ecosystem Processes in a Tropical Rainforest Forest Service researchers collaborated with external scientists to examine the effects of hurricane disturbance on tropical forest ecosystems. ...	2015
IITF-2013-025	Scientists Characterize Woody Debris to Assess Fuel Loads in Northeastern Puerto Rico Forest Service scientists characterized the amounts of woody debris (coarse woody debris, fine woody debris, duff, and litter) for eight differe ...	2013
IITF-2012-04	The Role of Millipedes in Tropical Ecosystems The direct and indirect effects on litter of varying lignin content	2012