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Fact Sheets
 FHTET Program Areas

Biological Control

Hemlock Wooly Adelgid

FHTET was created in 1995 by the Deputy Chief for State and Private Forestry to provide a focus within Forest Health Protection to develop and deliver technologies to protect and improve the health of America's forests. Biological control was identified as one of the major work emphasis areas for developing technologies to deal with the threat of invasive species to forests.

FHTET Biological Control Program

The FHTET Biological Control program (FHTET-BC) is tiered to the broader Forest Service's National Strategic Framework for Invasive Species Management as well as regional plans dealing with invasive species. FHTET-BC focuses its resources on a few insects and weed species that have the greatest potential for biological control.

The mission of the FHTET-BC is to demonstrate a strong leadership role in the development and implementation of biological control technologies to manage wide spread infestations of invasive species and to use biological control as a viable component for integrated pest-management efforts.

FHTET-BC program benefits resource managers by:

  • Identifying natural enemies for biological control of invasive species
  • Increasing awareness of new technologies and the implementation of biological control, through publications, workshops and scientific meetings and training sessions.
  • Coordinating and focusing funding for biological control.
  • Forming partnerships and coordinating the development and implementation of biological control technologies.
  • Developing recommendations for the restoration of native plant species

FHTET Biological Control Program: Sponsored Projects (3.4 MB PDF)
Overview of the Forest Health Technology Enterprise Team Biological Control Program for Invasive Species 1995 through 2017 (12 MB PDF)
A Global Compendium of Weeds

Biological Control of Invasive Native and Non-Native Plants (BCIP)

2018 BCIP Proposal Guidelines
BCIP Funded Proposals


FHTET-BC Program – 1995 through 1999
The initial five years (1995 through 1999) of the FHTET-BC Program were devoted to a comprehensive overview of the past history of biological control attempts for established invasive forest pests and documenting the recently introduced pests that were appropriate for biocontrol. These reviews were published as were the results of four sponsored meetings, all of which provided the initial basis for initiation of projects beyond 1999.

The extensive list of established invasive species introduced on forest and rangelands in the U.S. as well as potential non-native species required the FHTET-BC Program to focus its resources on a few priority insect and weed species that appeared to have the greatest potential for biological control. In the US, program staff focused on strengthening ongoing Biocontrol programs for arthropod species established in limited geographical areas (e.g. hemlock woolly adelgid and beech scale)  with the specific management objective of assisting in the search for and establishing additional natural enemies for each selected invasive species. Also, we were requested to provide assistance internationally in the development of biological control efforts for the pink hibiscus mealybug in Puerto Rico and Sirex woodwasp in several regions in Brazil. Because non-native invasive plants make up a large percentage of the species diversity in each state and numerous invasive species occupy the same site, species were selected for biological control based on their invasiveness. An effort was initiated to develop a biological control program for mile-a-minute, which is an invasive weed from China and Japan that was rapidly spreading in the Northeast.
The overseas exploration efforts to locate natural enemies and conduct host range tests were coordinated with Forest Service International Programs (Gary Man, Asian Pacific Coordinator) who provided many opportunities through financial support and contacts in China.

FHTET-BC – 2000 through 2004
The next five years (2000 through 2004) of the FHTET-BC Program focused on publishing documents, sponsoring meetings, continuing ongoing biological control projects, and initiating new projects to manage invasive species in forest ecosystems.

Major progress was made in searching for, locating, and rearing natural enemies of hemlock woolly adelgid in U.S. quarantine facilities, and then releasing and monitoring these natural enemies for establishment and impacts. Two species of coccinelids and a derodontid were released in the U.S. for control of hemlock woolly adelgid. These natural enemies were recovered from hemlock woolly adelgid populations in China, Japan, and British Columbia. Efforts to locate parasitoids of beech scale were not successful. Natural enemies of pink hibiscus mealybug and Sirex woodwasp were released and established in Puerto Rico and Brazil, respectively. Classical biological control efforts were initiated for emerald ash borer, ambermarked birch leafminer, and elongate hemlock scale.
Classical biological control efforts were initiated for non-native invasive plants: kudzu, Japanese knotweed, garlic mustard, and tree-of-heaven. A weevil from China, Rhinoncomimus latipes that feeds on mile-a-minute was approved for release in the U.S. in 2003; releases of this weevil were made in NJ and DE in 2004.
International Programs continued to provide much needed financial assistance in support of foreign exploration and host range testing.

FHTET-BC Program – 2005 through 2015
The program focus continued on publishing documents, sponsoring meetings, and continuing biological control projects. The biological control efforts were initiated for goldspotted oak borer, Chinese privet, and Japanese stiltgrass.

In 2010, as part of the national request for proposals (RFP) conducted by the Forest Service, Forest Health Protection, a request for technology development proposals for biological control of native and non-native plants (BCIP) was announced. The focus of the RFP was to develop new technologies in the use of biological controls in the fight against invasive plants. This RFP process continued on a yearly basis, with the FHTET Biological Control Program Manager and a multi-regional review team selecting proposals for funding. A total of 42 proposals were funded through 2015.

Two symposia on "Biological Control of Invasive Species of Forests" were co-sponsored by FHTET: 2007 in Beijing, China (FHTET-2008-04) and 2010 in Amherst, Massachusetts (Biocontrol 57[2]-113-348).

Three symposia, "International Symposium on Biological Control of Arthropods," were co-sponsored by FHTET: 2002, FHTET 2003-05; FHTET: 2005, FHTET-2005-08; FHTET: 2009, FHTET-2008-06.

Biological Control publications

About Biological Control

Biological control – the reduction of an organism's population density through use of its natural enemies – has been recognized as being one of the most effective and cost-efficient long-term approaches for managing invasive species. Natural enemies (parasites, predators, herbivores, and pathogens) reduce the population of hosts; in turn, host abundance influences the population levels of natural enemies.

"Classical" biological control is the intentional introduction of non-native natural enemies for permanent establishment and long-term control of invasive species in the infested areas. It is a strategy that has been used extensively to control non-native invasive species. It is a long-term process: successful classical biological control programs tend to average 10 years from the discovery and evaluation of natural enemies to their release and establishment in an infested area. Successful weed Biocontrol programs begin with localized damage to individual plants and finish with a sustained regional reduction in the plant population.

Program Manager

Richard Reardon

USDA Forest Service - FHTET
180 Canfield Street
Morgantown, WV 26505
Phone: (304) 285-1566
Fax: (304) 285-1564


Richard Reardon received a B.S. in Wildlife/Fisheries Management from Utah State University and a M.S. and Ph.D in Forest Entomology from the University of Michigan. He worked for the USDA Animal and Plant Health Inspection Service (APHIS) and Forest Service (Research and State and Private Forestry) to develop biopesticides and biological controls for agricultural and forest pests. Dick joined the Forest Health Technology Enterprise Team in 1995. | USFS | Forest Health Protection | FOIA | Accessibility Statement | Privacy Policy | Non-Discrimination Statement | Information Quality
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