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SPECIES:  Sequoiadendron giganteum
Visitor standing by a fire-scarred giant sequoia. Creative Commons image by Paul Bolstad, University of Minnesota, Bugwood.org.

 


Introductory

SPECIES: Sequoiadendron giganteum
AUTHORSHIP AND CITATION : Habeck, R. J. 1992. Sequoiadendron giganteum. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: https://www.fs.fed.us/database/feis/plants/tree/seqgig/all.html []. ABBREVIATION : SEQGIG SYNONYMS : Sequoia gigantea (Lindl.) Buchholz NRCS PLANT CODE : SEGI2 COMMON NAMES : giant sequoia bigtree TAXONOMY : The scientific name of giant sequoia is Sequoiadendron giganteum (Lindl.) Buchholz [26,28]. There are no recognized subspecies, varieties, or forms. LIFE FORM : Tree FEDERAL LEGAL STATUS : See OTHER STATUS OTHER STATUS : NO-ENTRY


DISTRIBUTION AND OCCURRENCE

SPECIES: Sequoiadendron giganteum
GENERAL DISTRIBUTION : The natural distribution of giant sequoia is restricted to about 75 groves, comprising a total area of only 35,607 acres (14,416 ha) along a limited area of the western Sierra Nevada, California.  The northern two-thirds of its range, from the American River in Placer County southward to the Kings River has only eight disjunct groves.  The remaining groves are concentrated between the Kings River and the Deer Creek Grove in southern Tulare County [10,28].  Groves range in size from approximately 2,470 acres (1,000 ha) with 20,000 giant sequoias to small groves with only six living trees [24].
Giant sequoia distribution. Map courtesy of USDA, NRCS. 2018. The PLANTS Database. National Plant Data Team, Greensboro, NC [26].
ECOSYSTEMS : 
   FRES21  Ponderosa pine
   FRES23  Fir - spruce
   FRES28  Western hardwoods


STATES : 
     CA

BLM PHYSIOGRAPHIC REGIONS : 
   4  Sierra Mountains


KUCHLER PLANT ASSOCIATIONS : 
   K005  Mixed conifer forest


SAF COVER TYPES : 
   243  Sierra Nevada mixed conifer


SRM (RANGELAND) COVER TYPES : 
   none


HABITAT TYPES AND PLANT COMMUNITIES : 
Giant sequoia principally occurs in scattered groves.  Nowhere does it
grow in pure stands, although in a few small areas stands do approach a
pure condition [28].  Although the giant sequoia groves of the central
and southern Sierra Nevada represent only a specific mesic segregate of
typical white fir (Abies concolor) forest communities, these groves are
often given special community recognition.  Only giant sequoia is
restricted to the groves [24].

Typically, giant sequoia is found in a mixed conifer type dominated by
California white fir (A. concolor var. lowiana).  Characteristic
associates include sugar pine (Pinus lambertiana), Jeffrey pine (P.
jeffreyi), ponderosa pine (P. ponderosa), Douglas-fir (Pseudotsuga
menziesii), incense-cedar (Calocedrus decurrens), and California black
oak (Quercus kelloggii).  Shrub types include bush chinkapin
(Castanopsis sempervirens) and mountain whitethorn (Ceanothus
cordulatus) [10,14,28].

Giant sequoia as a dominant species in the following typings:

Terrestrial natural communities of California [29]
Montane and subalpine vegetation of the Sierra Nevada and Cascade Ranges [24]

MANAGEMENT CONSIDERATIONS

SPECIES: Sequoiadendron giganteum
WOOD PRODUCTS VALUE : Giant sequoia was cut commercially from the 1850s up to the mid-1950s. Young giant sequoia has favorable wood properties.  It is decay-resistant and used as dimensional lumber, veneer, and plywood [21].  Old growth has low tensile strength and brittleness, making it unsuitable for most structural purposes.  The most historically popular items milled from giant sequoia were fenceposts, grape stakes, shingles, novelties, patio furniture, and pencils [10]. IMPORTANCE TO LIVESTOCK AND WILDLIFE : Only a limited number of wildlife species utilize giant sequoia for food and shelter. Birds:  Over 30 bird species have been identified in giant sequoia groves.  A variety of foliage- and air-feeding birds occupy the upper canopy, while sapsuckers feed through the thin bark.  Cavity-nesters that use giant sequoia for nesting include white-headed woodpeckers and flickers, and an occasional perching bird such as a nuthutch. Mammals:  Common mammal associates include the deer mouse, chipmunk, shrew, gray squirrel, golden-mantled ground squirrel, mule deer, coyote, black bear, and various reptiles.  Reports of chipmunks using giant sequoia sawdust for cleansing baths have been noted.  The chickaree is especially noted for its relationship to giant sequoia.  Chickarees make the soft flesh of green giant sequoia cone scales a major food item.  An individual chickaree may cut and eat as many as 3,000 to 3,500 cones per year [10,12]. PALATABILITY : Deer browse on 4- and 5-year-old giant sequoia; however, it is generally considered low in palatability [10]. NUTRITIONAL VALUE : The mean caloric value of giant sequoia seeds is 4,738 calories per gram dry weight.  The outer portions of the cones provide 4,690 calories per gram dry weight [12]. COVER VALUE : Wildlife primarily use giant sequoia for cover.  Early in giant sequoia development, large mammals use dense stands as hiding and thermal cover. Mature trees are used to a limited extent by arboreal species such as birds, squirrels, and other small mammals [10]. VALUE FOR REHABILITATION OF DISTURBED SITES : NO-ENTRY OTHER USES AND VALUES : Giant sequoia is planted as an ornamental inside and outside of its native range.  It is also used for Christmas trees [28]. OTHER MANAGEMENT CONSIDERATIONS : Insects:  Insects do not seriously harm giant sequoias older than about 2 years.  Carpenter ants (Campanotus laevigatus) do not directly harm the trees, although they do create pathways for fungi [28].  A wood-boring beetle (Trachykele opulenta) may kill trees damaged by road cuts or the undercutting of stream banks.  The larvae of this beetle may girdle a giant sequoia by feeding on the inner bark.  The cerambycid beetle (Phymatodes nitidus) lays its larvae in green giant sequoia cones.  Other cone larvae predators are the gelechiid moth (Gelechia spp.) and lygaeid bug (Ischnorrhynchus resedae).  In all, 151 species of insects and 37 arachnids are known to be associated with the giant sequoia in that they use it to complete some part of their life cycle [12,19,28]. Disease:  At least nine fungi have been found associated with decayed giant sequoia wood.  The most prevalent fungi are Heterobasidion annosum, Armillaria mellea, Poria incrassata, and P. albipellucida. Diseases generally do not kill trees past the seedling stage directly, but rather by contributing to root or stem failure.  No other types of disease, including seedling disease, are known to be problems to giant sequoia [12,19,28]. Air-pollution creating acidic mists significantly reduce root growth of giant sequoia [25].  The development of facilities for human use, such as paved roads and buildings, can damage giant sequoia roots and hence slow growth [27].

BOTANICAL AND ECOLOGICAL CHARACTERISTICS

SPECIES: Sequoiadendron giganteum
GENERAL BOTANICAL CHARACTERISTICS : Giant sequoias grow to an average height of 250 to 275 feet (76-84 m) and 15 to 20 feet (5-7 m) d.b.h.  Record trees have been reported to be 310 feet (95 m) in height and 35 feet (11 m) d.b.h.  The leaves are awl-shaped, sessile, and persistent.  Seed cones are 2 to 3 inches (5-8 cm) long, serotinous, persistent, and may remain green up to 20 years. Bark is fibrous, furrowed, and may be 2 feet (0.6 m) thick at the base of the columnar trunk [6,10].  The oldest known giant sequoia based on ring count is 3,200 years old [10].
Giant sequoia cone and needles. Creative Commons image by Tom DeGomez, University of Arizona, Bugwood.org.
RAUNKIAER LIFE FORM : 
Phanerophyte


REGENERATION PROCESSES : 
Giant sequoia regenerates primarily by seed, although occasionally it
may reproduce naturally by vegetative methods.  Giant sequoias up to
about 20 years of age may produce stump sprouts subsequent to injury.
Giant sequoia of all ages may sprout from the bole when old branches are
lost to fire or breakage.  Cuttings from juvenile donors root quickly
and in high percentages (up to 94 percent) [10].

Flowering and fruiting:  Giant sequoia is monoecious; male and female
cone buds form during late summer.  Pollination takes place between the
middle of April and May.  Fertilization usually occurs in August when
the cones are nearly full-sized.  Embryos develop rapidly during the
next summer and reach maturity at the end of the second growing season.

Seed production and dissemination:  Young trees start to bear cones at
the age of 20 years.  Cones may remain attached to the tree for 8 to 12
years and much of the seed will be retained.  During the late summer,
however, some seed is shed when the cone scales shrink.  Most seeds are
liberated when the cone dries out and becomes detached.  Each cone yields
an average of 230 seeds.  The average number of cleaned seeds per pound
is approximately 81,000 (200,070/kg).  Stored giant sequoia seed remains
moderately viable for many years [5,10,28].  At any given time, a large
tree may be expected to have approximately 11,000 cones.  The upper part
of the crown of any mature giant sequoia invariably produces a greater
abundance of cones than its lower portions.

A mature giant sequoia has been estimated to disperse from 300,000 to
400,000 seeds per year.  Seed dispersal results from seed falling from
the tree-top, insect and rodent activity, or by cones falling to the
ground.  The winged seeds may be carried up to 600 feet (183 m) from the
parent tree.

Seedling development:  Giant sequoia seeds germinate best when totally
buried in disturbed mineral soil.  April, May, September, and October
temperatures are best for early development.  Soil moisture conditions
and seedling survival are generally better in spring than during any
other season.  Light conditions are generally best for growing at
one-half full sunlight.  Upon germination, the seedling stands 3/4 to 1
inch (1.9-2.5 cm) high, usually with four cotyledons.  By autumn,
seedlings have up to six branches and are 3 to 4 inches (8-10 cm) tall.
After the second year, the seedling attains a height of 8 to 12 inches
(20-30 cm) with a taproot penetrating to a depth of 10 to 15 inches
(25-38 cm) [28].

Growth and yield:  Giant sequoia is the worlds largest tree in terms of
total volume.  Beyond the seedling stage, giant sequoia unhindered by an
overstory continues to grow at the same rate as its competitors.  Yields
of second growth stands dominated by giant sequoia were found to equal
or slightly exceed those of second-growth mixed-conifer stands on the
same site.  Lower branches die fairly readily from shading, but trees
less than 100 years old retain most of their dead branches.  Boles of
mature trees generally are free of branches to a height of 98 to 148
feet (30-40 m) [28].


SITE CHARACTERISTICS : 
Low temperatures seem to be the limiting factor for giant sequoia at the
upper elevational limits of its range, as well as in areas with severe
winters where the species has been introduced.  Distribution of giant
sequoia at lower elevations appears to be restricted to sites with
available soil moisture throughout the summer drought period [24,28].

Climate:  Giant sequoia is found in a humid climate characterized by dry
summers.  Mean annual precipitation varies from 35 to 55 inches (88-138
cm).  Most precipitation comes in the form of snow between October and
April.  Mean annual snowfall ranges from 144 to 197 inches (360-493 cm),
and snow depths of 6.6 feet (2 m) or greater are common.  Mean daily
maximum temperatures for July are typically 75 to 84 degrees Fahrenheit
(24-29 deg C).  Mean minimum temperatures for January vary from 34 to 21
degrees Fahrenheit (1 to -6 deg C) [28].

Soils and topography:  Most giant sequoia groves are on granitic-based
residual and alluvial soils.  Some groves are on glacial outwash from
granite.  Other common parent materials include schistose, dioritic and
andesitic rocks.  Giant sequoia grows best in deep, well-drained sandy
loams.  It occurs with higher frequency on mesic sites, such as drainage
bottoms and meadow edges.  Soil pH ranges from 5.5 to 7.5, with an
average of about 6.5.  Long-term site occupancy develops soil of high
fertility, good base status, and low bulk density.  Except for its
moisture content, soil typically plays only a minor role in influencing
the distribution of the species [28].

Elevation:  Elevation of the giant sequoia groves generally range from
4,590 to 6,560 feet (1,400-2,000 m) in the north, and 5,580 to 7,050
(1,700-2,150 m) to the south.  The lowest natural occurrence of the
species is 2,720 feet (830 m) and the highest is 8,860 feet (2,700 m).
Giant sequoia generally appears on southern slopes in its northern
distribution and on more northerly slopes in the south [28].


SUCCESSIONAL STATUS : 
Giant sequoia has adapted to keep its crown higher than that of its
associates.  On disturbed sites, giant sequoia is a strong competitor,
although never totally dominating a stand [10].  Current data does not
indicate that any enlargement of giant sequoia groves is taking place.
Mature giant sequoia mark the outer boundaries, which have remained
stable over a period of 500 to 1,000+ years.  High levels of
reproduction are not necessary to maintain the present population
levels.  Few groves, however, have sufficient young trees to maintain
the present density of mature giant sequoias for the future.  The
majority of giant sequoias are currently undergoing a gradual decline in
density since the European settlement days [24].


SEASONAL DEVELOPMENT : 
Giant sequoia flowers from April to May; cone ripening and seed
dispersal occurs in the spring and summer months.  Seeds dropped just
before the first snow or just as the snow melts may have the best chance
of germinating and becoming successfully established.  Growth of giant
sequoia generally begins in the early spring to late fall [28].

FIRE ECOLOGY

SPECIES: Sequoiadendron giganteum
FIRE ECOLOGY OR ADAPTATIONS : Fire is the most serious damaging agent to young giant sequoia. Seedlings and saplings are highly susceptible to mortality or serious injury by fire.  Giant sequoia exhibits the following adaptations to fire:  rapid growth, fire resistant bark, elevated canopies and self-pruned lower branches, latent buds, and serotinous cones [10,12]. Mature giant sequoia are more resistant to fire damage and few are killed by fire alone [28]. Giant sequoia groves represent a fire climax community whose stability is maintained by frequent fires.  In the absence of regular ground fires, litter accumulates on the forest floor and limits germination and establishment of seedlings [24].  Giant sequoia in Whitaker's Forest, California, produced 9,089 pounds per acre (10,181 kg per ha) of ground litter [3].  If these conditions are maintained in the future, the groves will become a long-standing seral community trending toward a mature white fir forest without giant sequoia [24]. FIRE REGIMES : Find fire regime information for the plant communities in which this species may occur by entering the species name in the FEIS home page under "Find Fire Regimes". POSTFIRE REGENERATION STRATEGY :    crown-stored residual colonizer; long-viability seed in on-site cones    off-site colonizer; seed carried by wind; postfire years 1 and 2    off-site colonizer; seed carried by animals or water; postfire yr 1&2

FIRE EFFECTS

SPECIES: Sequoiadendron giganteum
IMMEDIATE FIRE EFFECT ON PLANT : In Sequoia and Kings Canyon National Parks, a moderate-severity prescribed fire contributed little to the mortality of giant sequoia that were larger than 1 foot (0.30 m) d.b.h.  Additionally, there is no evidence that previous fire scarring had any relationship to tree mortality [5].  Low- to moderate-severity fires scorch the bark of giant sequoia and usually cause scarring.  High-severity fires may reach the crown and consume part or all of the canopy cover [2].  A direct relationship exists between the size of the basal fire scar in mature giant sequoias and the the likelihood of damage to the top or foliage of the trees [23].  Reduction of supporting wood from scarring predisposes the tree to falling, and provides an opening for fungi responsible for root disease and heart rot [28].
Prescribed fire in a giant sequoia grove in Kings Canyon National Park. US Park Service image.
DISCUSSION AND QUALIFICATION OF FIRE EFFECT : 
Despite the general belief that giant sequoia wood is not especially
flammable, it burns hotly when splintered and dry [10].


PLANT RESPONSE TO FIRE : 
High-severity fires will generally kill pole-size and younger trees.
Immediately following the passage of fire, seeds will drop as a reaction
to hot convectional air movement through the canopy.  Seeds will
germinate on the favorable mineral seedbeds created by the fire [5].

Postfire seedling establishment:  When high-severity fires burn in dense
stands of mature giant sequoias, as many as 40,485 seedlings per acre
(100,000/ha) may develop following heat-induced seedfall [11].  After a
prescribed burn in Sequoia and Kings Canyon National Parks, a
high-severity burn resulted in 40,000 seedlings per acre (98,800/ha) the
first year after burning.  A lower-severity burn resulted in 13,000
seedlings per acre (32,110/ha).  Not a single giant sequoia seedling was
found on the unburned control plot in this study [5].
DISCUSSION AND QUALIFICATION OF PLANT RESPONSE : 
A fall prescribed fire in the Tharp Creek Watershed of Sequoia National Park
resulted in no giant sequoia mortality on a white fir-mixed conifer
site monitored for 5 years after fire. The fire burned from 23 to 26 October
1990. Relative humidity during the day was 21% to 30% and at night was 30%
to 40%. Fuel moisture levels in the litter and duff averaged 28%. For 100-hour
and 1,000-hour fuels, moisture levels were 14% and 64%, respectively. At the
time of ignition, air temperatures were 50 to 61 ?F (10-16 ?C), and winds were
calm. The fire was a combination of backing and strip headfires with flame 
lengths of 0.16 to 7.9 feet (0.05-2.4 m). One-hour, 10-hour, and 100-hour fuels
were reduced by 96%, 77%, and 60%, respectively.  Tree (≥4.6 feet (1.4 m))
mortality was evaluated before and after fire as well as from an unburned
reference site. Basal area changes were also monitored before and after
the fire. Compared to the unburned control, mean annual percent change in giant
sequoia basal area increased by an average of 1.27% on the burned site before
the fire. From 1989 to 1994 (includes 1 year of prefire data), giant sequoia basal
area increased by 0.90% on the burned site compared to the control [31]. For
more information, see the entire Research Paper by Mutch and Parsons [31].

Summer prescribed fires (19 August-11 September 1969) promoted giant sequoia 
seedling establishment in a giant sequoia-mixed-conifer forest in Kings Canyon 
National Park.  The summer fires burned at moderate to high severities.  They 
also reduced the density of white fir and incense-cedar realative to giant 
sequoia and sugar pine.  Favorable establishment of giant sequoia seedlings 
was related to both parent tree density and fire severity. Giant sequoia 
seedling densities exceeded 40,000/acre (99,000/ha) severly burned plots. 
Kilgore [30] suggested that under similar fire weather conditions, follow-up 
prescribed fire would be needed 7 to 10 years later.  For further information, 
see the Research Project Summary of Kilgore and others' study.

FIRE MANAGEMENT CONSIDERATIONS : 
Restoration of historical stand structure and fire regimes is need if giant 
sequoia and other mixed-conifer forests are to continue through millennia to come. 
Research on the importance of periodic fire in maintaining natural giant
sequoia forests has justified the need to restore a natural fire regime.
The principal goal of fire management in giant sequoia groves in
Sequoia-Kings Canyon and Yosemite National Parks is to restore or
maintain the natural fire regime to the maximum extent possible.
Prescribed burns are now conducted by igniting fires in a spot pattern
and allowing nature to produce a mosaic of effects [20].

The long-standing fire suppression policy of federal and state land
agencies has created at least two major problems for the giant sequoia:
(1) the continuing reproduction of the species has been seriously
hampered and (2) the build-up of dead fuels and the growth of other
young trees in the understory pose threats of destructive forest fires
in the crowns of existing groves.  In 1969, the National Park Service
began a program or prescribed burning in Kings Canyon National Park.
Prescribed burning has produced relatively few deleterious side effects
on giant sequoia groves [27].

Prescribed burning is currently an active management strategy in giant
sequoia groves.  Fire prepares seedbeds, recycles nutrients, maintains
successional diversity, decreases the number of trees susceptible to
attack by insects and disease, reduces fire hazards, and favors wildlife
[5,10,27].  A prescribed burn in Kings Canyon National Park resulted in
an increase in flycatcher and robin numbers [4,13].  A number of changes
in bird and mammal populations are forecasted if fire is reintroduced on
a large scale.  High-severity fire will increase the number of
trunk-feeding birds preying on the increased amount of insects [12].

Prior to protection under Park status in 1864, the Mariposa Grove and
Yosemite National Park sustained fires every 20 to 25 years [9].  Other
research found that in presettlement times, any given site in the middle
elevations of the Sierra was burned over every 5 to 10 years [18,27].

REFERENCES

SPECIES: Sequoiadendron giganteum
REFERENCES :  1.  Bernard, Stephen R.; Brown, Kenneth F. 1977. Distribution of mammals,        reptiles, and amphibians by BLM physiographic regions and A.W. Kuchler's        associations for the eleven western states. Tech. Note 301. Denver, CO:        U.S. Department of the Interior, Bureau of Land Management. 169 p.        [434]  2.  Biswell, H. H. 1961. The big trees and fire. National Parks Magazine.        April: 1-4.  [8786]  3.  Biswell, H. H.; Gibbens, R. P.; Buchanan, H. 1966. Litter production by        bigtrees and associated species. California Agriculture. 20(9): 5-7.        [12692]  4.  Bock, Carl E.; Lynch, James F. 1970. Breeding bird populations of burned        and unburned conifer forest in the Sierra Nevada. Condor. 72: 182-189.        [5113]  5.  Boe, Kenneth N. 1974. Sequoiadendron giganteum (Lindl.) Buchholz Giant        sequoia. In: Schopmeyer, C. S., ed. Seeds of woody plants in the United        States. Agriculture Handbook No. 450. Washington: U. S. Department of        Agriculture, Forest Service: 767-768.  [7751]  6.  Brockman, C. Frank. 1979. Trees of North America. New York: Golden        Press. 280 p.  [16867]  7.  Eyre, F. H., ed. 1980. Forest cover types of the United States and        Canada. Washington, DC: Society of American Foresters. 148 p.  [905]  8.  Garrison, George A.; Bjugstad, Ardell J.; Duncan, Don A.; [and others].        1977. Vegetation and environmental features of forest and range        ecosystems. Agric. Handb. 475. Washington, DC: U.S. Department of        Agriculture, Forest Service. 68 p.  [998]  9.  Hartesveldt, R. J.; Harvey, H. T. 1968. The fire ecology of Sequoia        regeneration. In: Proceedings, Tall Timbers fire ecology conference;        1967 November 9-10; Hoberg. No. 7. Tallahassee, FL: Tall Timbers        Research Station: 65-77.  [6384] 10.  Hartesveldt, Richard J.; Harvey, H. Thomas; Shellhammer, Howard S.;        Stecker, Ronald E. 1975. The sequoia of the Sierra Nevada. Washington,        DC: U.S. Department of the Interior, National Park Service. 180 p.        [4233] 11.  Harvey, H. Thomas; Shellhammer, Howard S. 1991. Survivorship and growth        of giant sequoia (Sequoiadendron giganteum (lindl.)buchh.) seedlings        after fire. Madrona. 38(1): 14-20.  [14879] 12.  Harvey, H. Thomas; Shellhammer, Howard S.; Stecker, Ronald E. 1980.        Giant sequoia ecology: Fire and reproduction. Scientific Monograph        Series No 12. Washington, DC: U.S. Department of the Interior, National        Park Service. 182 p.  [6587] 13.  Kilgore, Bruce M. 1971. Response of breeding bird populations to habitat        changes in a giant sequoia forest. American Midland Naturalist. 85(1):        135-152.  [7281] 14.  Kilgore, Bruce M. 1972. Fire's role in a Sequoia forest. Naturalist.        23(1): 26-37.  [8783] 15.  Kuchler, A. W. 1964. Manual to accompany the map of potential vegetation        of the conterminous United States. Special Publication No. 36. New York:        American Geographical Society. 77 p.  [1384] 16.  Lambert, Sherman; Stohlgren, Thomas J. 1988. Giant sequoia mortality in        burned and unburned stands. Journal of Forestry. 86(2): 44-46.  [7221] 17.  Lyon, L. Jack; Stickney, Peter F. 1976. Early vegetal succession        following large northern Rocky Mountain wildfires. In: Proceedings, Tall        Timbers fire ecology conference and Intermountain Fire Research Council        fire and land management symposium; 1974 October 8-10; Missoula, MT. No.        14. Tallahassee, FL: Tall Timbers Research Station: 355-373.  [1496] 18.  Martin, Robert E. 1982. Fire history and its role in succession. In:        Means, Joseph E., ed. Forest succession and stand development research        in the Northwest: Proceedings of a symposium; 1981 March 26; Corvallis,        OR. Corvallis, OR: Oregon State University, Forest Research Laboratory:        92-99.  [9830] 19.  Parmeter, John R., Jr. 1986. Diseases and insects of giant sequoia. In:        Weatherspoon, C. Phillip; Iwamoto, Y. Robert; Piirto, Douglas D.,        technical coordinators. Proceedings of the workshop on management of        giant sequoia; 1985 May 24-25; Reedley, CA. Gen. Tech. Rep. PSW-95.        Berkeley, CA: U.S. Department of Agriculture, Forest Service, Pacific        Southwest Forest and Range Experiment Station: 11-13.  [9803] 20.  Parsons, David J.; Nichols, H. Thomas. 1986. Management of giant sequoia        in the national parks of the Sierra Nevada, California. In:        Weatherspoon, C. Phillip; Iwamoto, Y. Robert; Piirto, Douglas D.,        technical coordinators. Proceedings of the workshop on management of        giant sequoia; 1985 May 24-25; Reedley, CA. Gen. Tech. Rep. PSW-95.        Berkeley, CA: U.S. Department of Agriculture, Forest Service, Pacific        Southwest Forest and Range Experiment Station: 26-29.  [9807] 21.  Piirto, Douglas D. 1986. Wood of giant sequoia: properties and unique        characteristics. In: Weatherspoon, C. Phillip; Iwamoto, Y. Robert;        Piirto, Douglas D., technical coordinators. Proceedings of the workshop        on management of giant sequoia; 1985 May 24-25; Reedley, CA. Gen. Tech.        Rep. PSW-95. Berkeley, CA: U.S. Department of Agriculture, Forest        Service, Pacific Southwest Forest and Range Experiment Station: 19-23.        [9806] 22.  Raunkiaer, C. 1934. The life forms of plants and statistical plant        geography. Oxford: Clarendon Press. 632 p.  [2843] 23.  Rundel, Philip W. 1973. The relationship between basal fire scars and        crown damage in giant sequoia. Ecology. 54(1): 210-213.  [6639] 24.  Rundel, Philip W.; Parsons, David J.; Gordon, Donald T. 1977. Montane        and subalpine vegetation of the Sierra Nevada and Cascade Ranges. In:        Barbour, Michael G.; Major, Jack, eds. Terrestrial vegetation of        California. New York: John Wiley & Sons: 559-599.  [4235] 25.  Temple, Patrick J. 1988. Injury and growth of Jeffrey pine and giant        sequoia in response to ozone and acidic mist. Environmental and        Experimental Botany. 28(4): 323-333.  [13016] 26.  USDA Natural Resources Conservation Service. 2017. PLANTS Database, [Online]. U.S. Department of Agriculture, Natural Resources Conservation Service (Producer). Available: https://plants.usda.gov/. [34262] 27.  Vale, Thomas R. 1975. Ecology and environmental issues of the Sierra        Redwood (Sequoiadendron giganteum), now restricted to California.        Environmental Conservation. 2(3): 179-188.  [8776] 28.  Weatherspoon, C. Philip. 1990. Sequoiadendron giganteum (Lindl.)        Buchholz. In: Burns, Russell M.; Honkala, Barbara H., technical        coordinators. Silvics of North America. Volume 1. Conifers. Agric.        Handb. 654. Washington, DC: U.S. Department of Agriculture, Forest        Service: 552-562.  [13415] 29.  Holland, Robert F. 1986. Preliminary descriptions of the terrestrial        natural communities of California. Sacramento, CA: California Department        of Fish and Game. 156 p.  [12756] 30.  Kilgore, B. M. 1973. Impact of prescribed burning on a Sequoia-mixed        conifer forest. In: Proceedings, annual Tall Timbers fire ecology conference; 1972 June        8-10; Lubbock, TX. No. 12. Tallahassee, FL: Tall Timbers Research Station:        345-375.  [6270] 31.  Mutch, Linda S.; Parsons, David J. 1998. Mixed conifer forest        mortality and establishment before and after prescribed fire in Sequoia        National Park, California. Forest Science. 44(3): 341-355.  [29033]

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