Index of Species Information
SPECIES: Ambrosia dumosa
SPECIES: Ambrosia dumosa
AUTHORSHIP AND CITATION:
Marshall, K. Anna. 1994. Ambrosia dumosa. In: Fire Effects Information System, [Online].
U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station,
Fire Sciences Laboratory (Producer). Available:
On 28 June 2018, the common name of this species was changed in FEIS
from: white bursage
to: burrobush. Images were also added.
Franseria dumosa Gray 
NRCS PLANT CODE:
The scientific name for burrobush is Ambrosia dumosa (Gray) Payne .
It is a member of the aster family (Asteraceae). There are no recognized
FEDERAL LEGAL STATUS:
No special status
DISTRIBUTION AND OCCURRENCE
SPECIES: Ambrosia dumosa
Burrobush occurs throughout the Sonoran and Mojave deserts although
it is typically considered a Mojave Desert species . It ranges north
to Death Valley, California, southern Nevada, and southwestern Utah. It
extends along the Gulf in Baja California as far south as Bajia Los
Angeles and into Sonora as far south as Tiburon Island .
|Distribution of burrobush in the United States. Map courtesy of USDA, NRCS. 2018. The PLANTS Database.
National Plant Data Team, Greensboro, NC [2018, June 28] .
FRES30 Desert shrub
AZ CA NV UT MEXICO
BLM PHYSIOGRAPHIC REGIONS:
3 Southern Pacific Border
6 Upper Basin and Range
7 Lower Basin and Range
12 Colorado Plateau
KUCHLER PLANT ASSOCIATIONS:
K042 Creosotebush - bursage
K043 Paloverde - cactus shrub
SAF COVER TYPES:
SRM (RANGELAND) COVER TYPES:
HABITAT TYPES AND PLANT COMMUNITIES:
Burrobush is a dominant or codominant member of most plant
communities in the Sonoran and Mojave deserts. It usually occurs in
open, species-poor communities with creosote bush (Larrea tridentata).
At the northern boundary of burrobush, in the transition zone
between the Mojave and Great Basin deserts, associated species of the
creosote bush-burrobush community include wolfberry (Lycium spp.),
range ratany (Krameria parvifolia), Mojave yucca (Yucca schidigera),
California jointfir (Ephedra funera), spiny hopsage (Grayia spinosa),
and winterfat (Krascheninnikovia lanata) [38,47]. The density of
burrobush is about 2,500 plants per hectare [3,47].
Approximately 70 percent of the Mojave Desert is covered with open or
very open stands of creosote bush and burrobush [15,29,40].
Associated species in the Mojave Desert include desertsenna (Cassia
armata), Nevada ephedra (Ephedra nevadensis), white burrobrush
(Hymenoclea salsola), and wolfberry .
In the Sonoran Desert, associated members of the creosote bush-
burrobush community are acacia (Acacia paucipina), fourwing saltbush
(Atriplex canescens), ocotillo (Fouquieria splendens), big galleta
(Hilaria rigida), cholla (Opuntia spp.) and western honey mesquite
(Prosopis glandulosa var. torreyana) . In the Arizona Upland
Subdivision of the Sonoran Desert, the density of burrobush is 549.7
plants per hectare and burrobush cover is 2.7 percent. In the Lower
Colorado River Valley, the density of burrobush is 84 plants per
hectare and burrobush cover is 0.1 percent .
In addition to the creosote bush-burrobush association, burrobush
is a member of the following associations: Joshua tree (Yucca
brevifolia)-big galleta , saguaro (Carnegiea gigantea)-paloverde
(Cercidium spp.) , Sonoran creosote bush scrub, Mojave creosote bush
scrub, and Mojave mixed woody scrub .
Publications listing burrobush as a dominant or codominant species
Sonoran Desert 
Preliminary descriptions of the terrestrial natural communities of California
Vegetation of the Santa Catalina Mountains: community types and
Mojave Desert scrub vegetation 
SPECIES: Ambrosia dumosa
IMPORTANCE TO LIVESTOCK AND WILDLIFE:
Burrobush is an important browse species in several areas of the
Sonoran Desert. Browsing pressure on burrobush is particularly
heavy during years of low precipitation, when production of winter
annuals is low .
Burrobush is of intermediate forage value . It is fair to good
forage for horses and fair to poor for cattle and sheep. However,
because there is often little other forage where burrobush grows, it
is often highly valuable to browsing animals . Webb  observed
that sheep browsed primarily on new growth and seeds.
In the Mojave Desert, 8 percent of mature burrobush plants were
browsed by black-tailed jackrabbits. Forty-three percent of
transplanted seedlings were browsed. Fourteen percent of browsed
seedlings were more than 90 percent consumed .
Many desert rodents, including kangaroo rats, eat burrobush seeds
Burrobush is moderately palatable to cattle and sheep and slightly
more palatable to horses. Closely cropped burrobush plants on
heavily stocked range indicates inadequate forage rather than high
Burrobush is palatable to feral asses. Fecal analysis indicated
that burrobush was the primary forage used by feral asses in winter
The nutrient value of burrobush fluctuates seasonally; it is greater
in the spring and less in the fall . In a 2-year study in the Lower
Colorado River Valley, burrobush had a gross energy value of about
4.2 kilocalories per gram. Crude protein was highest in February at 10
percent, declining the rest of the year to 4 to 7 percent. Phosphorus
content was highest in spring at 1,110 milligrams per kilogram of plant
material, declining to 500 milligrams per kilogram in August and
In the northern Mojave Desert, Romney and others  estimated the
nutrient load of new burrobush leaves as follows:
Nitrogen 1.29 kg/ha
Phosphorous 0.114 kg/ha
Sodium 0.035 kg/ha
Potassium 1.70 kg/ha
Calcium 0.93 kg/ha
Magnesium 0.17 kg/ha
Reichman  estimated that burrobush seeds contain 3,838 calories
per gram or 23.72 calories per seed.
VALUE FOR REHABILITATION OF DISTURBED SITES:
Burrobush may be used to revegetate disturbed sites in southwestern
deserts. For instance, burrobush may be planted along California
highways where unirrigated perennial vegetation has not survived .
Burrobush may be planted from containerized plants with a high
probability of success. Plantings should be made in late winter or
early spring, although the time of planting is less important than the
vigor of the seedlings. Rodent protectors should be used .
OTHER USES AND VALUES:
Burrobush is a host for sandfood (Pholisma sonorae), a parasitic
plant with a sweet, succulent, subterranean flowerstalk. Sandfood was
a valuable food supply for desert peoples [58,59].
OTHER MANAGEMENT CONSIDERATIONS:
Creosote bush-burrobush communities are poorly suited for livestock
grazing because of low productivity and low water availability .
Burrobush is sensitive to browsing. Browsing significantly
decreased the cover and volume of burrobush by 27 and 21 percent,
respectively, in the Mojave Desert . In the Lower Colorado River
Valley, overbrowsing decreased the cover of burrobush from 2.26 to
0.04 percent .
Pollution from electric power generating facilities may also decrease
burrobush. Burrobush showed intermediate sensitivity to sulphur
dioxide and nitrogen dioxide fumigation .
BOTANICAL AND ECOLOGICAL CHARACTERISTICS
SPECIES: Ambrosia dumosa
GENERAL BOTANICAL CHARACTERISTICS:
Burrobush is a native, drought-deciduous rhizomatous shrub growing
from 8 to 24 inches (20-60 cm) tall. Its many slender, stiff branches
form a compact, hemispherical crown [14,41]. The leaves of
burrobush are small and deeply divided. They may become so dry that it is
difficult to tell whether they are alive or dead .
The staminate and pistillate heads of burrobush intermingle
throughout the length of its racemes . The pistillate heads are
two-flowered, producing obovoid fruits 0.2 to 0.8 inches (5-20 mm) long.
The burs generally contain 20-35 flattened, scattered, unhooked spines
that are about 0.08 inches (2 mm) long .
The root system of burrobush is derived from a segmented root crown,
and is mostly comprised of lateral roots . Roots may grow 5 to 15
times the length of the stem  and extend to a depth of 28 inches (70
cm) . Anderson  found the shoot to root dry weight ratio of
burrobush to be 1.38. The leaf to root ratio was 0.18.
Definitive information on the longevity of burrobush is not
available in the literature. Due to cloning, burrobush may be an
extremely long-lived shrub . Some researchers, however, have
suggested that the longevity of burrobush is similar to that of its
noncloning congener, triangle bursage (A. deltoidea): somewhat less
than 50 years [31,63].
RAUNKIAER LIFE FORM:
Burrobush reproduces both vegetatively and sexually.
Vegetative reproduction: Muller  and Wright and Howe  have
described vegetative reproduction in burrobush. As burrobush
plants age, their crowns open irregularly with the successive deaths of
individual aerial shoots. The clone slowly spreads to as much as 3.3
feet (1 m) in diameter. The original seedling shoot branches
intricately at its base and below the soil. The short rhizomes root
independently so that the death of the seedling stem and subsequent
rotting away of the original root crown disconnects the rhizomatous
shoots. Windblown soil and organic debris accumulate about the base of
burrobush clones, producing a sizable mound over time .
Sexual reproduction: Burrobush flowers anytime during the spring,
summer, and fall if enough rain falls . It produces seeds
abundantly, and seedlings establish in open space . Large numbers
of burrobush seedlings emerge following heavy fall precipitation
. In September of 1976, after a record rainfall near Ocotillo,
California, the density of burrobush seedlings was 466 plants per
acre (1,151 plants/ha) .
Burrobush seeds have prickles that easily enter and remain in skin
and hair, so burrobush is probably dispersed by mammals. Although
burrobush has moderately heavy fruits with low lofting ability,
requiring an air current of 87.9 centimeters per second, Maddox and
Carlquist  suggested that the tumbling ability of the fruits aids in
Germination experiments have been performed on burrobush. Young and
Young  found that 30 days of moist stratification treatment at 35
degrees Fahrenheit (1.7 deg C) markedly improved burrobush
germination. Graves and others  found that both moist sand
stratification and carbon treatments improved 7- and 14-day germination
Burrobush commonly grows on arroyos, bajadas, gentle slopes, valley
floors, and sand dunes at elevations up to 3,000 feet (900 m) throughout
the Sonoran and Mojave deserts [27,32,41,44,53]. It occurs on
calcareous, sandy, alluvial soil that is often underlain by a caliche
hardpan [3,13,29,41,54]. Burrobush grows in pure stands or with
associates, especially creosote bush, in barren or open areas [24,31,41].
Temperatures in the Sonoran and Mojave deserts are variable and extreme.
At Puerto Libertad, Sonora, near the southern boundary of burrobush
distribution, the mean annual temperature is 68.37 degrees Fahrenheit
(20.2 deg C). Daytime temperatures in the summer often reach 117
degrees Fahrenheit (47 deg C) . In Rock Valley, Nevada, near the
northern boundary of burrobush distribution, temperatures range from
5 degrees Fahrenheit (-15 deg C) in winter to 117 degrees Fahrenheit (47
deg C) in summer .
Phenological events in the Sonoran and Mojave deserts are triggered by
rain. In the Sonoran, rainfall averages 4 to 12 inches (100-300 mm)
annually with a bimodal distribution . The Mojave gets more winter
than summer rain . In Rock Valley, Nevada, rainfall averages 5.524
inches (138.1 mm), with 60 percent falling between September and February
Low soil oxygen may be a controlling factor in the distribution of
desert species. Burrobush was more tolerant of low soil oxygenation
than creosote bush .
Succession in the desert is difficult to characterize because there is
no clear change in species composition over time. For instance, in
creosote bush-burrobush communities, both burrobush and
creosote bush persist in the community even though changes in their
relative abundances may occur.
Most burrobush are located on bare soil away from other plants.
McAuliffe  found that 83 to 92 percent of all young burrobush in
creosote bush-burrobush communities were located in bare spaces.
Burrobush was the principal colonizer of open spaces in those
Once established, burrobush acts as a nurse plant to creosote bush
and other desert species, providing improved microhabitat and protection
from herbivory [16,31]. McAuliffe  found that 85.5 percent of all
young creosote bush were either rooted beneath the canopies of live
burrobush or positioned next to dead ones. Most creosote bush
establishment apparently occurs while the burrobush are alive. The
smallest creosote bush in McAuliffe's study were associated exclusively
with live burrobush.
Because of its colonizing ability, burrobush is a common pioneer on
disturbed areas in the Mojave Desert . In a comparison between
vegetation in disturbed and undisturbed sites, burrobush was
subdominant to creosote bush on control sites and dominant on disturbed
sites 40 years after disturbance . Vasek  noted that while
burrobush colonizes open space by large-scale seedling
establishment, creosote bush depends upon cloning or requires
burrobush for establishment.
Creosote bush-burrobush communities are probably adapted to continual
or relatively slight disturbance such as lightly shifting sand surfaces
[49,50]. If slight disturbance does occur, creosote bush-burrobush
communities recover quickly in terms of species composition .
Burrobush leafs out in February or March; the mean leafing-out date
in Rock Valley, Nevada, was February 23. The mean flowering date was
May 2 . Burrobush seeds usually germinate following heavy
September precipitation . A minimum amount of rainfall is required
to induce germination. For instance, a 1971 rain of 1 to 1.96 inches
(25-49 mm) was sufficient but neither an August 1972 rain of 0.68 inch
(17 mm) nor a July rain of 0.84 inch (21 mm) promoted germination .
Burrobush has a drought dormancy period in the summer and may have
an induced dormancy period during the winter if freezing night
temperatures kill its leaves .
SPECIES: Ambrosia dumosa
FIRE ECOLOGY OR ADAPTATIONS:
There is little mention of fire in relation to burrobush in the
literature. One study described limited sprouting and seedling
establishment after fire .
Fires in the desert are infrequent and of low severity because
production of annual and perennial herbs seldom provides a fuel load
capable of sustaining fire. In the Mojave Desert, there is little
record of fires. Humphrey  stated that the creosote bush-burrobush
community is "essentially nonflammable" because the shrubs are too
sparse to carry fire.
Woody remains of burrobush take about 40 years to decay beyond the
point of recognitions .
POSTFIRE REGENERATION STRATEGY:
Small shrub, adventitious-bud root crown
Rhizomatous shrub, rhizome in soil
Initial-offsite colonizer (off-site, initial community)
Find fire regime information for the plant communities in which this
species may occur by entering the species name in the FEIS home page under
"Find Fire Regimes".
SPECIES: Ambrosia dumosa
IMMEDIATE FIRE EFFECT ON PLANT:
Fire generally kills burrobush. A low-severity fire occurred in the
Coachella Valley, California, after 7 years of above normal
precipitation. Some burrobush plants survived because the fire
burned patchily. However, most burrobush plants burned because
their canopies contained numerous small branches in proximity to
herbaceous fuels. A few burrobush plants were only scorched; they
retained most of their branches and dried foliage. Eighty-nine percent
of burrobush plants were killed by the fire .
DISCUSSION AND QUALIFICATION OF FIRE EFFECT:
PLANT RESPONSE TO FIRE:
Burrobush sprouted at low rates after the low-severity fire in the
Coachella Valley, California. Only 6 out of 16 scorched plants and 2
out of 81 burned plants sprouted . Burrobush seedling
establishment on burned sites was poor during the first growing season
after the fire but increased in later growing seasons. Poor seedling
establishment was probably unrelated to seed availability since
burrobush occurred in adjacent unburned areas . In postfire year 5,
cover of burrobush on burned sites was 1 percent while cover on
adjacent unburned sites was about 5.8 percent .
DISCUSSION AND QUALIFICATION OF PLANT RESPONSE:
The Research Project Summary Nonnative annual grass fuels and fire in
California's Mojave Desert provides information on prescribed fire and
postfire response of plant community species, including burrobush,
that was not available when this species review was written.
FIRE MANAGEMENT CONSIDERATIONS:
Biomass production and thus fuel loadings vary seasonally and annually
for burrobush. In 1971 and 1974 in Rock Valley, Nevada, estimated
abovegound stem dry mass of burrobush was 31 and 41 grams per square
meter, respectively. Net aboveground production was consistently higher
in spring than in fall. In 1973, after an above average year of
rainfall, flowers and fruits made up almost half of the new tissue
produced by burrobush .
SPECIES: Ambrosia dumosa
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