SPECIES: Osmunda cinnamomea
SPECIES: Osmunda cinnamomea
AUTHORSHIP AND CITATION :
Walsh, Roberta A. 1994. Osmunda cinnamomea. In: Fire Effects Information System, [Online].
U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station,
Fire Sciences Laboratory (Producer). Available:
SCS PLANT CODE :
COMMON NAMES :
The currently accepted scientific name of cinnamon fern is Osmunda
cinnamomea L. It is in the family Osmundaceae [7,26]. Recognized
subspecies, varieties, and form are as follows:
O. c. ssp. asiatica (Fern.) Hulten (found in eastern Asia) 
O. c. var. cinnamomea [22,52]
O. c. var. glandulosa Waters [9,18,52]
O. c. forma frondosa (T. & G.) Britt. [7,49,52,62]
LIFE FORM :
Fern or Fern Ally
FEDERAL LEGAL STATUS :
No special status
OTHER STATUS :
DISTRIBUTION AND OCCURRENCE
SPECIES: Osmunda cinnamomea
GENERAL DISTRIBUTION :
Cinnamon fern occurs in North America from Newfoundland to western
Ontario and south to the Gulf States and New Mexico. It also occurs in
eastern Asia [7,22].
FRES10 White - red - jack pine
FRES11 Spruce - fir
FRES12 Longleaf - slash pine
FRES13 Loblolly - shortleaf pine
FRES14 Oak - pine
FRES15 Oak - hickory
FRES16 Oak - gum - cypress
FRES18 Maple - beech - birch
FRES19 Aspen - birch
FRES41 Wet grasslands
AL AR CT DC FL GA IL IN KY LA
ME MD MA MI MN MS NH NJ NM NY
NC OH PA RI SC TN TX VT VA WV
WI NB NF NS ON PE PQ
BLM PHYSIOGRAPHIC REGIONS :
KUCHLER PLANT ASSOCIATIONS :
K073 Northern cordgrass prairie
K079 Palmetto prairie
K091 Cypress savanna
K093 Great Lakes spruce - fir forest
K094 Conifer bog
K095 Great Lakes pine forest
K096 Northeastern spruce - fir forest
K097 Southeastern spruce - fir forest
K100 Oak - hickory forest
K102 Beech - maple forest
K103 Mixed mesophytic forest
K104 Appalachian oak forest
K106 Northern hardwoods
K107 Northern hardwoods - fir forest
K108 Northern hardwoods - spruce forest
K110 Northeastern oak - pine forest
K111 Oak - hickory - pine forest
K112 Southern mixed forest
K113 Southern floodplain forest
SAF COVER TYPES :
12 Black spruce
13 Black spruce - tamarack
17 Pin cherry
18 Paper birch
19 Gray birch - red maple
20 White pine - northern red oak - red maple
21 Eastern white pine
22 White pine - hemlock
23 Eastern hemlock
24 Hemlock - yellow birch
37 Northern white-cedar
52 White oak - black oak - northern red oak
70 Longleaf pine
74 Cabbage palmetto
80 Loblolly pine - shortleaf pine
81 Loblolly pine
82 Loblolly pine - hardwood
83 Longleaf pine - slash pine
97 Atlantic white-cedar
98 Pond pine
104 Sweetbay - swamp tupelo - redbay
108 Red maple
SRM (RANGELAND) COVER TYPES :
HABITAT TYPES AND PLANT COMMUNITIES :
Cinnamon fern is listed as a habitat type or indicator species in:
Freshwater Wetlands: A guide to common indicator plants of the northeast 
Application of forest habitat-type classification system in Michigan and
Species associated with cinnamon fern are listed for bogs in
northeastern Illinois , southern New Hampshire , North Carolina
[28,46,53,61], Louisiana [2,36], and eastern Texas ; peatlands in
northcentral Minnesota ; Florida gulf coast hydric hammocks ,
cypress heads [16,45], and bayheads ; pocosins in several states ;
South Carolina wet longleaf pine savannahs ; and an eastern hemlock
(Tsuga canadensis)-fern community with mesic soil and high humidity in
northeastern West Virginia .
SPECIES: Osmunda cinnamomea
IMPORTANCE TO LIVESTOCK AND WILDLIFE :
Cinnamon fern was grazed by cattle in southeastern North Carolina in
pond pine (Pinus rigida var. serotina) forests. It ranked second only
to cane (Arundinaria gigantea) in cattle preference. In May, before
cane fully leafed out, 30 to 50 percent of available cinnamon fern
herbage was utilized, after which fern utilization practically ceased.
Cinnamon fern was palatable for about a month after fronds unrolled .
White-tailed deer were observed grazing substantial amounts of cinnamon
fern in southwestern Virginia in 1982 .
NUTRITIONAL VALUE :
COVER VALUE :
VALUE FOR REHABILITATION OF DISTURBED SITES :
OTHER USES AND VALUES :
Cinnamon fern coiled fiddlehead leaves up to 8 inches (20 cm) tall can
be collected in the spring, steamed or boiled, and eaten .
Cinnamon fern spore germination in liquid medium is useful for bioassay
of toxic copper, cadmium, and zinc concentrations .
OTHER MANAGEMENT CONSIDERATIONS :
Cattle grazed cinnamon fern in southeastern North Carolina, but because
of the short time it was utilized, grazing had relatively little effect
on the total fern stand. In places where grazing was heavy, the density
and vigor of cinnamon fern was noticeably reduced. Cinnamon fern was
more abundant on unlogged than on logged sites, in both grazed and
ungrazed conditions .
Cinnamon fern in southeastern Connecticut was an associate in lowland
hardwood and shrub communities subjected to 20 years of herbicide use on
trees to maintain shrubs. Cinnamon fern cover was 1 to 5 percent both
before herbicide treatment began in 1953 and after 20 years of treatment .
Cinnamon fern in Atlantic white-cedar (Chamaecyparis thyoides) wetlands
in New Jersey occurred in sites of all disturbance classes studied
including undisturbed sites, those in housing developments, and those at
stormwater drain outfalls .
Cinnamon fern in eastern Quebec was present in a northern hardwoods site
that was clearcut and subjected to three experimental disturbance
treatments: prepared with a V-blade (high intensity), prepared with a
toothed brush rake (medium intensity), or disked (low intensity).
Cinnamon fern was more common on the low-disturbance site but survived
on other sites .
Cinnamon fern in naturally regenerated, mature slash pine (Pinus
elliottii) flatwoods in southeastern Florida was present at 1.3
kilograms per hectare foliage biomass. The site was then clearcut in
the fall of 1978, prepared by burning, shearing and piling, discing and
bedding, and planted to slash pine in 1979. In two subsequent
vegetation surveys in the summers of 1980 and 1981, cinnamon fern was
not present .
Cinnamon fern frequently forms large clumps  and may produce almost
all the understory cover in swamps with dense overstory shade .
BOTANICAL AND ECOLOGICAL CHARACTERISTICS
SPECIES: Osmunda cinnamomea
GENERAL BOTANICAL CHARACTERISTICS :
Cinnamon fern is a perennial fern which is native to the eastern United
States. Rhizomes of sporophytic plants are stout, woody , and
creeping  to suberect ; the roots are fibrous . Sporophytes
have separate fertile and sterile pinnate fronds are covered with thick
hairs when immature. Some hairs are still present on fertile fronds at
maturity . Sterile fronds are up to 6 feet (1.8 m) long  and 6
to 12 inches (15-30 cm)  wide. Fertile fronds are shorter than
sterile fronds, and the pinnae are much smaller, nonphotosynthetic, and
cinnamon brown. Sporangia are clustered, naked, large, globose, and
bivalved . Cinnamon fern spores are green, with functional
chloroplasts. The spores germinate into photosynthetic, platelike,
thalloid gametophytes [23,42].
RAUNKIAER LIFE FORM :
REGENERATION PROCESSES :
Cinnamon fern spores have very short viability after release from the
sporophyte. They either fail to germinate or germinate poorly after
just a few weeks . Under controlled conditions, spores germinate at
high percentages at 41 degrees Fahrenheit (5 deg C), and they also
germinate at higher temperatures .
One study showed that cinnamon fern sporophytes allelopathically
inhibited germination of cinnamon fern spores . Another study did
not demonstrate this effect .
SITE CHARACTERISTICS :
Cinnamon fern is found on poorly drained low ground and in thickets, wet
marshy woods , swamps, ditches, and streambanks . It is
generally found in ombrotrophic bogs , but it also grows on
minerotrophic wooded island hummocks in peatlands . It is usually
associated with sphagnum (Sphagnum spp.)  in wet acid soils with
high organic content [28,46,55]; it is an indicator of such soils in the
Haut-Saint-Laurent region, Quebec . Where humidity is very high
cinnamon fern can sometimes be found on better drained soils .
Cinnamon fern has been reported at the following elevations:
Elevation (feet) Elevation (m)
Florida 125-141 38-43 
Louisiana 197-276 60-84 
Maryland 0-51 0-16 
New York 210-3,124 64-952 
North Carolina 39-2,917 12-889 [28,60,64]
South Carolina 45-75 14-23 
West Virginia 1,096-2,625 334-800 [6,39]
Ontario 581 177 
SUCCESSIONAL STATUS :
Facultative Seral Species
Cinnamon fern is considered a late seral species in the bog seres of the
northern United States and Laurentian Canada, but may not persist to
climax stages . In the Adirondack upland flora cinnamon fern is
intolerant to midtolerant of shade . Cinnamon fern in west
Louisiana occurs in bogs that are mostly open, with a few scattered
trees and shrubs . However, cinnamon fern occurs in heavy shade
under a closed canopy along the Gulf Coast of Florida . It also
occurs under shade in Atlantic white-cedar wetlands in New Jersey 
and in baldcypress (Taxodium distichum) swamps in eastern Maryland .
Cinnamon fern in a southern Ontario bog was subjected to disturbance by
peat mining which removed all vegetation and up to 6.6 feet (2 m) of
peat. The mined areas had been abandoned to natural, unassisted
regeneration for 1, 6, 10, and 24 years. Cinnamon fern occurred in all
disturbance classes, but did not occur in the undisturbed control site
SEASONAL DEVELOPMENT :
Both sterile and fertile cinnamon fern fronds expand during a short
period in early spring [32,37]. Leaf expansion is complete within about
a month. Fertile fronds begin to wither in early summer, after
sporulation is completed . Sterile pinnae begin to wither at the
end of summer, and the stipe somewhat later, until the entire aerial
part of the plant is dry . Cinnamon fern spores are discharged in
spring. They can germinate within 1 or 2 days of release .
Cinnamon fern sporulates from March through July, depending on latitude
SPECIES: Osmunda cinnamomea
FIRE ECOLOGY OR ADAPTATIONS :
Although fronds are probably killed by fire during the growing season,
cinnamon fern sprouts from rhizomes after the aerial portions are burned
[49,53]. Cinnamon fern has good fire tolerance and shows vigorous
rhizome growth after fire . Spores germinate on mineral soil ,
so cinnamon fern probably colonizes after fire. It grows in the open
conditions created by fire in at least part of its range [20,36,60].
Cinnamon fern occurs in the Greater Sandhills of south-central North
Carolina in Atlantic white-cedar dominated wetland corridors. The trees
in these wetlands show evidence of past fires . Cinnamon fern
occurs in the Green Swamp, North Carolina, longleaf pine/wiregrass
savannahs which are maintained by frequent fire . It also occurs in
fire-maintained South Carolina longleaf pine savannahs .
Find fire regime information for the plant communities in which this
species may occur by entering the species name in the FEIS home page
under "Find Fire Regimes".
POSTFIRE REGENERATION STRATEGY :
Rhizomatous herb, rhizome in soil
Secondary colonizer - off-site seed
SPECIES: Osmunda cinnamomea
IMMEDIATE FIRE EFFECT ON PLANT :
Cinnamon fern fronds are probably killed by fire.
DISCUSSION AND QUALIFICATION OF FIRE EFFECT :
PLANT RESPONSE TO FIRE :
Cinnamon fern appears to be well adapted to recurring fires. It
occurred in a west Louisiana bog that was burned on a regular basis
during the winter . It also occurred in a southern Mississippi peat
bog burned annually for 7 years . Cinnamon fern occurs in North
Carolina Coastal Plain longleaf pine-wiregrass savannahs that are fire
maintained. The savannahs have fire intervals typically less than 8
years . Cinnamon fern in the wetter areas of a virgin longleaf pine
forest in southwestern Georgia increased greatly over 27 years of annual
burning. In this area cinnamon fern disappears within about 5 years in
the absence of fire, due in part to competition from shrubs and young
Cinnamon fern occurred on a fire barren in southwestern Nova Scotia that
had burned 2 years before. It was thought to be a component of the
prefire vegetation that survived the fire .
Cinnamon fern occurs in both burned and unburned tree islands in the
Florida Everglades, but is much more abundant on unburned islands. Fire
on these islands tends to burn out the peat substrate during periods of
DISCUSSION AND QUALIFICATION OF PLANT RESPONSE :
Cinnamon fern usually increases slightly in response to fire.
Cinnamon fern in an eastern white pine (Pinus strobus)-mixed hardwood
forest in New Hampshire was subjected to October 1976 and April 1977
prescribed fires. Cinnamon fern occurred throughout the forest in moist
pockets and depressions. Fires were of low severity, with flames 10 to
18 inches (25.4-45.7 cm) high and scorch heights 4 to 6 feet (1.2-1.8
m). Cinnamon fern increased in cover and importance value after the
fall fire, but not after the spring fire or on control plots. Cinnamon
fern also occurred sporadically in adjacent eastern white pine forest
plots that were burned at the same times. Fire severity was less, with
flames 4 to 6 inches (10.2-15.2 cm) high and scorch heights 3 to 4 feet
(0.9-1.2 m). Cinnamon fern was listed as neutral in response to fire on
these sites .
FIRE MANAGEMENT CONSIDERATIONS :
Cinnamon fern survives fires in many habitat types and may increase in
cover . However, if the fire is so intense or of such duration that
it burns the organic substrate completely, cinnamon fern will be lost .
Cinnamon fern occurs on hillside seepage bogs in longleaf pine savannah
of east-central Texas that is burned every 3 to 5 years during the
nongrowing season to maintain savannah vegetation .
Cinnamon fern in bayheads of south Florida may survive wet-season fires,
but drought-season fires can destroy them by burning out the islands .
Cinnamon fern occurs in North Carolina Coastal Plain shrub bog
communities that become grass (Poaceae)-sedge (Cyperaceae) bogs when
In northeastern Minnesota the summer moisture content of 21 groups of
understory plants was evaluated for fire prediction purposes from June
24, 1976 (after the period of primary plant growth) to August 26, 1976.
Ferns, including cinnamon fern, were evaluated as a group. Fern
moisture was approximately in the middle range when compared to other
SPECIES: Osmunda cinnamomea
1. Abrahamson, Warren G.; Johnson, Ann F.; Layne, James N.; Peroni, Paricia
A. 1984. Vegetation of the Archbold Biological Station, Florida: an
example of the Southern Lake Wales Ridge. Florida Scientist. 47(4):
2. Allen, Charles M.; Stagg, Charles H.; Parris, Stephen D. 1988. Analysis
of the vegetation in pitcher plant bogs in two baygalls at Ft. Polk in
west central Louisiana. The Proceedings of the Louisiana Academy of
Sciences. 50: 1-6. 
3. Ash, A. N.; McDonald, C. B.; Kane, E. S.; Pories, C. A. 1983. Natural
and modified pocosins: literature synthesis and management options.
FWS/OBS-83/04. Washington, DC: U.S. Fish and Wildlife Service, Division
of Biological Sciences. 156 p. 
4. Beaven, George Francis; Oosting, Henry J. 1939. Pocomoke Swamp: a study
of a cypress swamp on the eastern shore of Maryland. Bulletin of the
Torrey Botanical Club. 66: 376-389. 
5. Bernard, Stephen R.; Brown, Kenneth F. 1977. Distribution of mammals,
reptiles, and amphibians by BLM physiographic regions and A.W. Kuchler's
associations for the eleven western states. Tech. Note 301. Denver, CO:
U.S. Department of the Interior, Bureau of Land Management. 169 p.
6. Bush, Eleanor M. 1988. A floristic study of a wet meadow in Barbour
County, West Virginia. Castanea. 53(2): 132-139. 
7. Cody, William J.; Britton, Donald M. 1989. Ferns and fern allies of
Canada. Ottawa, ON: Agriculture Canada, Research Branch. 430 p. 
8. Conde, Louis F.; Swindel, Benee F.; Smith, Joel E. 1983. Plant species
cover, frequency, and biomass: Early responses to clearcutting,
chopping, and bedding in Pinus elliottii flatwoods. Forest Ecology and
Management. 6: 307-317. 
9. Crandall, Dorothy L. 1965. County distribution of ferns and fern allies
in Rhode Island. American Fern Journal. 55(3): 98-112. 
10. Dansereau, Pierre; Segadas-Vianna, Fernando. 1952. Ecological study of
the peat bogs of eastern North America. Canadian Journal of Botany.
30(5): 490-520. 
11. Dittberner, Phillip L.; Olson, Michael R. 1983. The plant information
network (PIN) data base: Colorado, Montana, North Dakota, Utah, and
Wyoming. FWS/OBS-83/86. Washington, DC: U.S. Department of the Interior,
Fish and Wildlife Service. 786 p. 
12. Dunlop, D. A. 1987. Community classification of the vascular vegetation
of a New Hampshire peatland. Rhodora. 89(860): 415-440. 
13. Ehrenfeld, Joan G.; Schneider, John P. 1991. Chamaecyparis thyoides
wetlands and suburbanization: effects on hydrology, water quality and
plant community composition. Journal of Applied Ecology. 28(2): 467-490.
14. Eleuterius, L. N.; Jones, S. B., Jr. 1969. A floristic and ecological
study of pitcher plant bogs in south Mississippi. Rhodora. 71: 29-34.
15. Elias, Thomas S.; Dykeman, Peter A. 1982. Field guide to North American
edible wild plants. [Place of publication unknown]
: Outdoor Life Books.
286 p. 
16. Ewel, Katherine Carter. 1984. Effects of fire and wastewater on
understory vegetation in cypress domes. In: Ewel, Katherine Carter;
Odum, Howard T., eds. Cypress swamps. Gainesville, FL: University of
Florida Press: 119-126. 
17. Eyre, F. H., ed. 1980. Forest cover types of the United States and
Canada. Washington, DC: Society of American Foresters. 148 p. 
18. Fernald, Merritt Lyndon. 1950. Gray's manual of botany. [Corrections
supplied by R. C. Rollins]. Portland, OR: Dioscorides Press. 1632 p.
(Dudley, Theodore R., gen. ed.; Biosystematics, Floristic & Phylogeny
Series; vol. 2). 
19. Francis , Patrick C.; Petersen, Raymond L. 1983. Eff. of copper,
cadmium, & zinc on percent spore germ. of the cinnamon fern (Osmunda
cinnamomea) and the sensitive fern (Onoclea sensibilis). Bulletin of
Environmental Contamination and Toxicology. 30(5): 559-566. 
20. Gaddy, L. L. 1982. The floristics of three South Carolina pine
savannahs. Castenea. 47: 393-402. 
21. Garrison, George A.; Bjugstad, Ardell J.; Duncan, Don A.; [and others]
1977. Vegetation and environmental features of forest and range
ecosystems. Agric. Handb. 475. Washington, DC: U.S. Department of
Agriculture, Forest Service. 68 p. 
22. Gleason, Henry A.; Cronquist, Arthur. 1991. Manual of vascular plants of
northeastern United States and adjacent Canada. 2nd ed. New York: New
York Botanical Garden. 910 p. 
23. Hill, R. H.; Wagner, W. H. 1974. Seasonality and spore type of the
Pteridophytes of Michigan. Michigan Botanist. 13: 40-44. 
24. Jobidon, Robert. 1990. Short-term effect of 3 mechanical site
preparation methods on species diversity. Tree Planters' Notes. 41(4):
25. Jonsson-Ninniss, Susan; Middleton, John. 1991. Effect of peat extraction
on the vegetation in Wainfleet Bog, Ontario. Canadian Field-Naturalist.
105(4): 505-511. 
26. Kartesz, John T.; Kartesz, Rosemarie. 1980. A synonymized checklist of
the vascular flora of the United States, Canada, and Greenland. Volume
II: The biota of North America. Chapel Hill, NC: The University of North
Carolina Press; in confederation with Anne H. Lindsey and C. Richie
Bell, North Carolina Botanical Garden. 500 p. 
27. Kirkman, W. Benson; Wentworth, Thomas R.; Ballington, James R. 1989. The
ecology and phytosociology of the creeping blueberries, Vaccinium
section Herpothamnus. Bulletin of the Torrey Botanical Club. 116(2):
28. Kologiski, Russell L. 1977. The phytosociology of the Green Swamp, North
Carolina. Tech. Bull. No. 250. Raleigh, NC: North Carolina Agricultural
Experiment Station. 101 p. 
29. Komarek, E. V., Sr. 1973. Ancient fires. In: Proceedings, annual Tall
Timbers fire ecology conference; 1972 June 8-9; Lubbock, TX. Number 12.
Tallahassee, FL: Tall Timbers Research Station: 219-240. 
30. Kotar, J. 1986. Application of forest habitat-type classification system
in Michigan and Wisconsin. In: Site classification in relation to forest
management: Proceedings of a symposium; 1985 August 27-29; Sault Ste.
Marie, ON. COJFRC Symposium Proceedings O-P-14. [Place of publication
: Canadian Forestry Service, Great Lakes Forestry Centre: 47-52.
31. Kuchler, A. W. 1964. Manual to accompany the map of potential vegetation
of the conterminous United States. Special Publication No. 36. New York:
American Geographical Society. 77 p. 
32. Kudish, Michael. 1992. Adirondack upland flora: an ecological
perspective. Saranac, NY: The Chauncy Press. 320 p. 
33. Lemieux, G. J. 1963. Soil-vegetation relationships in northern hardwoods
of Quebec. In: Forest-soil relationships in North America. Corvallis,
OR: Oregon State University Press: 163-176. 
34. Loomis, Robert M.; Roussopoulos, Peter J.; Blank, Richard W. 1979.
Summer moisture contents of understory vegetation in northeastern
Minnesota. Res. Pap. NC-179. St. Paul, MN: U.S. Department of
Agriculture, Forest Service, North Central Forest Experiment Station. 7
35. Loveless, Charles M. 1959. A study of the vegetation in the Florida
Everglades. Ecology. 40(1): 1-9. 
36. MacRoberts, B. R.; MacRoberts, M. H. 1988. Floristic composition of two
west Louisiana pitcher plant pogs. Phytologia. 65(3): 184-190. 
37. Maeda, Osamu. 1970. On the dry matter product. of two ferns, Osmunda
cinnamomea & Dryopteris crassirhizoma, in relation to their geographical
distribution in Japan. Japanese Journal of Botany. 20(3): 237-267.
38. Magee, Dennis W. 1981. Freshwater wetlands: A guide to common indicator
plants of the Northeast. Amherst, MA: University of Massachusetts Press.
245 p. 
39. Maguire, D. A.; Forman, R. T. 1983. Herb cover effects on tree seedling
patterns in a mature hemlock-hardwood forest. Ecology. 64(6): 1367-1380.
40. Martin, J. Lynton. 1956. An ecological survey of burned-over forest land
in southwestern Nova Scotia. Forestry Chronicle. 32: 313-336. 
41. Meilleur, A.; Bouchard, A.; Bergeron, Y. 1992. The use of understory
spp. as indicators of landform ecosystem type in heavily disturb.
forest: an evaluat. in the Haut-Saint-Laurent, Quebec. Vegetatio. 102:
42. Miller, J. H. 1968. Fern gametophytes as experimental material.
Botanical Review. 34(4): 361-440. 
43. Mohlenbrock, Robert H. 1986. (Revised edition). Guide to the vascular
flora of Illinois. Carbondale, IL: Southern Illinois University Press.
507 p. 
44. Mohlenbrock, Robert H. 1992. Boykin Springs Longleaf, Texas. Natural
History. July: 62-65. 
45. Monk, Carl D.; Brown, Timothy W. 1965. Ecological consideration of
cypress heads in north-central Florida. American Midland Naturalist. 74:
46. Moore, Julie H.; Carter, J. H., III. 1987. Habitats of white cedar in
North Carolina. In: Laderman, Aimlee D., ed. Atlantic white cedar
wetlands. [Place of publication unknown]: Westview Press: 177-190.
47. Niering, William A.; Goodwin, Richard H. 1974. Creation of relatively
stable shrublands with herbicides: arresting "succession" on
rights-of-way and pastureland. Ecology. 55: 784-795. 
48. Petersen, Raymond L.; Fairbrothers, David E. 1980. Reciprocal allelpathy
between the gametophytes of Osmunda cinnamomea and Dryopteris
intermedia. American Fern Journal. 70(2): 73-78. 
49. Radford, Albert E.; Ahles, Harry E.; Bell, C. Ritchie. 1968. Manual of
the vascular flora of the Carolinas. Chapel Hill, NC: The University of
North Carolina Press. 1183 p. 
50. Raunkiaer, C. 1934. The life forms of plants and statistical plant
geography. Oxford: Clarendon Press. 632 p. 
51. Ross, S. Rachel. 1978. The effects of prescribed burning on ground cover
vegetation of white pine and mixed hardwood forests in southeastern New
Hampshire. Durham, NH: University of New Hamshire. 151 p. Thesis.
52. Seymour, Frank Conkling. 1982. The flora of New England. 2d ed.
Phytologia Memoirs 5. Plainfield, NJ: Harold N. Moldenke and Alma L.
Moldenke. 611 p. 
53. Shepherd, W. O.; Dillard, E. U.; Lucas, H. L. 1951. Grazing and fire
influences in pond pine forests. Tech. Bull. No. 97. Raleigh, NC: North
Carolina State College, Agricultural Experiment Station. 56 p. In
cooperation with: U.S. Department of Agriculture, Forest Service,
Southeastern Forest Experiment Station. 
54. Stickney, Peter F. 1989. Seral origin of species originating in northern
Rocky Mountain forests. Unpublished draft on file at: U.S. Department of
Agriculture, Forest Service, Intermountain Research Station, Fire
Sciences Laboratory, Missoula, MT; RWU 4403 files. 7 p. 
55. Taft, John B.; Solecki, Mary Kay. 1990. Vascular flora of the wetland
and prairie communities of Gavin Bog and Prairie Nature Preserve, Lake
County, Illinois. Rhodora. 92(871): 142-165. 
56. U.S. Department of Agriculture, Soil Conservation Service. 1982.
National list of scientific plant names. Vol. 1. List of plant names.
SCS-TP-159. Washington, DC. 416 p. 
57. Vince, Susan W.; Humphrey, Stephen R.; Simons, Robert W. 1989. The
ecology of hydric hammocks: A community profile. Biological Rep.
85(7.26). Washington, DC: U.S. Department of the Interior, Fish and
Wildlife Service, Research and Development. 82 p. 
58. Wade, Dale; Ewel, John; Hofstetter, Ronald. 1980. Fire in south Florida
ecosystems. Gen. Tech. Rep. SE-17. Asheville, NC: U.S. Department of
Agriculture, Forest Service, Southeastern Forest Experiment Station. 125
59. Wagner, Holliday B.; Long, Karen E. 1991. Allelopathic effects of
Osmunda cinnamomea on three species of Dryopteris. American Fern
Journal. 81(4): 134-138. 
60. Walker, Joan; Peet, Robert K. 1983. Composition and species diversity of
pine-wiregrass savannas of the Green Swamp, North Carolina. Vegetatio.
55: 163-179. 
61. Wells, B. W. 1928. Plant communities of the Coastal Plain of North
Carolina and their successional relations. Ecology. 9(2): 230-242.
62. Werth, Charles R.; Haskins, Melanie L.; Hulburt, Akke. 1985. Osmunda
cinnamomea forma frondosa at Mountain Lake, Virginia. American Fern
Journal. 75(4): 128-132. 
63. Wheeler, Gerald A.; Glaser, Paul H.; Gorham, Eville; [and others]. 1983.
Contributions to the Red Lake peatland, northern Minnesota, with special
attention to Carex. American Midland Naturalist. 110(1): 62-96. 
64. Pittillo, J. Dan; Wagner, W. H., Jr.; Farrar, Donald R.; Leonard, S. W.
1975. New Pteridophyte records in the Highlands Biological Station area,
southern Appalachians. Castanea. 40(4): 263-272. 
65. Sauitis, Eva; Freeman, Deborah. [n.d.]. An illustrated guide to the
ferns of the Mackie Camp. Unpublished report. Fredonia, NY: State
University of New York, Biology Department. 56 p. 
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