Index of Species Information
WILDLIFE SPECIES: Megascops asio
WILDLIFE SPECIES: Megascops asio
AUTHORSHIP AND CITATION :
Sullivan, Janet. 1995. Megascops asio. In: Fire Effects Information System, [Online].
U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station,
Fire Sciences Laboratory (Producer). Available:
Otus asio (Linnaeus)
COMMON NAMES :
little horned owl
The currently accepted scientific name of eastern screech-owl is Megascops asio .
It is a member of the family Strigidae [27,41]. It was recently split from western screech-owl (M. kennicottii) which was formerly treated as a subspecies [3,21].
These two species interbreed in the Big Bend region of Texas, where their ranges overlap .
Marshall  accepted five subspecies:
M. a. maxwelliae (Ridgway), Rocky Mountain screech-owl
M. a. hasbroucki (Ridgway), Hasbrouck's screech-owl
M. a. mccalli (Cassin), Texas screech-owl
M. a. asio (Linneaus), southern screech-owl
M. a. floridanus (Ridgway), Florida screech-owl
FEDERAL LEGAL STATUS :
See OTHER STATUS
OTHER STATUS :
The eastern screech-owl is listed as a species of special concern on the
American Ornithologists' Union Blue List. It was first blue-listed in
1981, and was listed again in 1982 and 1986. Numbers of eastern
screech-owls have been reported as declining on the Hudson-Delaware,
southern Atlantic Coast, and in Appalachian and mid-western prairie
WILDLIFE DISTRIBUTION AND OCCURRENCE
WILDLIFE SPECIES: Megascops asio
GENERAL DISTRIBUTION :
The eastern screech-owl is found in eastern North America and
northeastern Mexico. Its range extends across southern Saskatchewan,
southern Manitoba, northern Minnesota, northern Michigan, southern
Ontario, southwestern Quebec, and Maine; south through eastern Montana,
eastern Colorado, and Texas to central Nuevo Leon, eastern San Luis
Potosi, and southern Tamaulipas; east to the Gulf Coast and southern
Florida [1,27]. At the western edges of its range, it appears to be
confined by the eastern front ranges of the Rocky Mountains in Wyoming,
Montana, and Colorado [1,13,22].
Distribution of subspecies is as follows:
Rocky Mountain screech-owl: Southeastern Saskatchewan, southern
Manitoba, eastern Montana, and the Dakotas south to eastern Wyoming,
western Nebraska, western Kansas, and northeastern Colorado. Possibly
also breeds in central Alberta, but these birds may be western screech-owl.
Hasbrouck's screech-owl: Central Kansas to Oklahoma and Texas.
Texas screech-owl: Lower Rio Grande to the southern border to Tamaulipas.
Southern screech-owl: Minnesota, peninsular Michigan, southern
Quebec, and southern Maine south to Missouri and northern parts of
Mississippi, Alabama, and Georgia.
Florida screech-owl: Florida and the Gulf Coast west at least to
Louisiana and north to Arkansas .
FRES10 White-red-jack pine
FRES12 Longleaf-slash pine
FRES13 Loblolly-shortleaf pine
BLM PHYSIOGRAPHIC REGIONS :
13 Rocky Mountain Piedmont
14 Great Plains
15 Black Hills Uplift
16 Upper Missouri Basin and Broken Lands
KUCHLER PLANT ASSOCIATIONS :
K081 Oak savanna
K082 Mosaic of K074 and K100
K084 Cross Timbers
K089 Black Belt
K093 Great Lakes spruce-fir forest
K094 Conifer bog
K095 Great Lakes pine forest
K096 Northeastern spruce-fir forest
K097 Southeastern spruce-fir forest
K098 Northern floodplain forest
K099 Maple-basswood forest
K100 Oak-hickory forest
K101 Elm-ash forest
K102 Beech-maple forest
K103 Mixed mesophytic forest
K104 Appalachian oak forest
K106 Northern hardwoods
K107 Northern hardwoods-fir forest
K108 Northern hardwoods-spruce forest
K110 Northeastern oak-pine forest
K111 Oak-hickory-pine forest
K112 Southern mixed forest
K113 Southern floodplain forest
K115 Sand pine scrub
SAF COVER TYPES :
1 Jack pine
5 Balsam fir
12 Black spruce
14 Northern pin oak
15 Red pine
20 White pine-northern red oak-red maple
21 Eastern white pine
22 White pine-hemlock
23 Eastern hemlock
24 Hemlock-yellow birch
25 Sugar maple-beech-yellow birch
26 Sugar maple-basswood
27 Sugar maple
28 Black cherry-maple
39 Black ash-American elm-red maple
40 Post oak-blackjack oak
42 Bur oak
43 Bear oak
44 Chestnut oak
45 Pitch pine
46 Eastern redcedar
50 Black locust
51 White pine-chestnut oak
52 White oak-black oak-northern red oak
53 White oak
55 Northern red oak
58 Yellow-poplar-eastern hemlock
59 Yellow-poplar-white oak-northern red oak
60 Beech-sugar maple
61 River birch-sycamore
65 Pin oak-sweetgum
72 Southern scrub oak
88 Willow oak-water oak-diamondleaf oak
89 Live oak
93 Sugarberry-American elm-green ash
94 Sycamore-sweetgum-American elm
95 Black willow
96 Overcup oak-water hickory
108 Red maple
110 Black oak
SRM (RANGELAND) COVER TYPES :
810 Longleaf pine-turkey oak hills
811 South Florida flatwoods
812 North Florida flatwoods
813 Cutthroat seeps
820 Everglades flatwoods
PLANT COMMUNITIES :
The eastern screech-owl occupies a wide variety of habitat types .
It is commonly associated with open woodlands of oaks (Quercus spp.),
maples (Acer spp.), and hickories (Carya spp.); it is also found in pine
(Pinus spp.) forests and plantations, mixed woodlands, orchards, and
forested wetlands . At the northern and western edges of its range,
the eastern screech-owl occupies riparian woods dominated by hardwoods
including boxelder (Acer negundo), green ash (Fraxinus pennsylvanica),
and bur oak (Quercus macrocarpa) . In South Dakota eastern
screech-owls were found in riparian woods dominated by eastern
cottonwood (Populus deltoides) and willows (Salix spp.), hardwoods
dominated by green ash, boxelder, American elm, and bur oak, and
shelterbelts. Siberian elm (Ulmus pumila) is a common shelterbelt tree
in South Dakota and is used by eastern screech-owls . In Kansas
eastern screech-owls were also observed in shelterbelts . In New
England, eastern screech-owls are found in aspen (Populus spp.), paper
birch (Betula papyrifera), northern hardwoods, red maple (Acer rubrum),
balsam fir (Abies balsamea), eastern hemlock (Tsuga canadensis),
northern red oak (Quercus rubra), eastern white pine (Pinus
strobus)-northern red oak-red maple, and eastern white pine communities
. In South Carolina on the Coastal Plain, eastern screech-owls were
observed in pine and hardwood stands. The managed pine stands were
mostly longleaf pine (P. palustris) and loblolly pine (P. taeda).
Hardwood stands were dominated by gums (Nyssa spp.), baldcypress
(Taxodium distichum), oaks, or red maple . In Louisiana eastern
screech-owls were found in mature mixed bottomland hardwoods dominated
by water oak (Q. nigra), pignut hickory (Carya glabra), American elm
(Ulmus americana), and sugarberry (Celtis laevigata). They also
inhabited cottonwood plantations and riverfront hardwoods dominated by
sweet pecan (Carya illinoiensis), water hickory (C. aquatica),
sugarberry, and waterlocust (Gleditsia aquatica) .
BIOLOGICAL DATA AND HABITAT REQUIREMENTS
WILDLIFE SPECIES: Megascops asio
TIMING OF MAJOR LIFE HISTORY EVENTS :
Diurnal Activity: Eastern screech-owls are rarely active before dusk.
In Ithaca, New York, the earliest recorded time of initiation of feeding
activity was 8:25 p.m.; the latest time feeding ended was 4:15 a.m. .
Migration Status: Eastern screech-owls are resident throughout their
range; suggestions that some migration occurs at the northern edge of
the range have not been substantiated . The large majority of adult
birds (87%) were recovered within 10 miles (16 km) of the banding site.
None had moved more than 40 miles (64 km) .
Pair Bond: Craighead and Craighead  stated that they had no data on
year-round pairing, but they observed birds that appeared to remain
loosely paired over the winter, using the same hunting areas and
roosting sites as during the breeding season. They delimited the
beginning of the breeding season by the selection of a nesting
territory; in Michigan this usually occurs around the end of February
Nesting: Peak nesting season, when young are in the nest, appears to
coincide with most passerine migration (a rich source of prey) . In
the Southeast, nesting season is from mid-March to late May, peaking in
April . In Michigan earliest egg date was April 18 (1942) . In
New York and New England, egg dates ranged from April 12 to May 18. In
Pennsylvania and New Jersey eggs were present in nests from March 23 to
May 19, and in Florida, eggs were present from March 11 to May 18 .
The shortest time between eggs is 48 hours; the time between eggs is
often 72 hours and occasionally longer . Typical clutch sizes are 3
to 5 eggs; clutch sizes range from 1 to 10 .
Incubation: Incubation is almost exclusively by the female; if she
leaves the nest to drink or bathe, the male will incubate until she
returns and pushes him away. The male brings food but the female eats
very little during incubation . The incubation period is variable,
lasting from 26 to 32 days .
Development of Young: The altricial young are downy, with eyes closed
. The young have been observed shivering almost constantly for the
first 2 weeks of life . The eyes are usually open by the 20th day.
Flight feathers begin coming in around the 25th day. The average time
in the nest is 4 to 5 weeks [7,19]. Fledglings stay on branches near
the nest until full flight is achieved. Even after leaving the nest
tree, fledglings continue to be fed by the parents for up to 5 weeks
after fledging. By the ninth week, fledglings usually begin molting to
the first-winter plumage. In Ohio the overall nesting success rate
(the proportion of active nests that fledged at least one young) was
86.1 percent, and the overall breeding success rate (the proportion of
eggs that resulted in young fledged) was 73.8 percent .
Juvenile Dispersal: Family groups begin to split up by late August.
Dispersal, in an apparently random direction, occurs in early fall
[18,19]. In a Connecticut banding study, juvenile dispersal fell into
two groups; juveniles either travelled fewer than 20 miles (32 km) or
much greater distances (a minority). The longest distance recorded for
juvenile dispersal was 185 miles (300 km) .
Breeding Age and Longevity: One banding study indicated that at least
25 percent of eastern screech-owls bred as yearlings . In the wild,
the average lifespan for eastern screech-owl in central Texas was
estimated as 3.6 years, although some individuals survived to at least 8
years of age . Captive birds have lived more than 20 years .
PREFERRED HABITAT :
Suitable eastern screech-owl habitat seems to be wherever nesting and
roosting cavities are near or adjacent to open areas with available
rodent prey . Eastern screech-owls usually inhabit open woodlands
adjacent to meadows, marshes, or fields; forest clearings; old orchards;
and parks and other open areas in urban or suburban areas [10,9]. They
are more tolerant of human habitation and human activities than other
owls, and are found in or near barns, outbuildings, birdhouses,
hedgerows, and fencerows [19,29]. They are rarely found in swamps or
deep forest habitats .
Eastern screech-owls generally prefer areas with widely spaced trees
interspersed with grassy openings . In Connecticut, red maple
woodlands, upland woodland, evergreen hedgerows, and edge habitats were
used by eastern screech-owl more often than expected if habitat choice
were random. Lawns, mixed woods, and evergreen woods were used less
often than would be expected if habitat choice were random. Although
lawns were not a selected habitat, they were a major component of all
monthly home ranges (a monthly home range is the amount of territory
used in the span of a month) . In central Kentucky radio-tagged
eastern screech-owls used woodlots and edges more than expected based on
availability, and used pastures, old fields, and croplands less than
expected . Lynch and Smith  reported that eastern screech-owl
abundance in Connecticut was positively related to the amount of natural
area within urban open space (examples include shrubs, old fields, and
marshes). There was also a positive relationship among eastern
screech-owl abundance, total habitat diversity, and the amount of linear
edge. Areas with mixtures of habitats including relatively high amounts
of undisturbed successional communities and edges tended to support many
owls . In the southeastern states, eastern screech-owls are found
up to 4,500 feet (1,371 m) elevation (but are rare at the higher
Hunting: Eastern screech-owls usually hunt in grassy openings, fields,
meadows, and along wooded field margins and streams .
Home Range: In suburban areas most food gathering trips are less than
330 feet (100 m) round trip. In areas with lower prey density food
trips may be longer . Home range size is related to prey
availability. Smith and Gilbert  estimated that the area traversed
in one night (the nightly range) for female eastern screech-owls was the
largest (38 acres [15.5 ha]) from November to February (when prey was
most scarce), was smallest during egg-laying and incubation (13 acres
[5.4 ha]) in March and April, and expanded again in June to 22 acres
(8.9 ha). The larger nightly ranges in winter were thought to reflect a
need to cover greater areas to obtain adequate food. The total home
range of one female was 323 acres (130.9 ha), but she typically hunted
only a small area each night and a larger cumulative portion each month.
Home range was largest while a female was selecting a nest site and
smallest during egg laying, incubation, and care of young. The actual
defended territory included only the area immediately around the nest
cavity . Gehlbach  estimated Connecticut home ranges as 9.9 to
14.8 acres (4-6 ha) and Texas home ranges exceeding 74.1 acres (30 ha).
Estimated Population Density: In the Southeast, sampled breeding
density was listed as 2.8 pairs per 100 acres (2.8 pairs/40 ha) .
Lynch and Smith  observed a range of 1 to 18 eastern screech-owls
per square mile (1-7/sq km); an average of 6 birds per square mile
(2.3/sq km) occurred in 4 suburban areas in Connecticut.
COVER REQUIREMENTS :
Nesting: Eastern screech-owls nest in natural cavities, woodpecker
holes, nest boxes, and other artificial structures . Within the
oak-hickory forest region, maples, apples (Malus spp.), sycamores
(Platanus occidentalis) with natural cavities, and pines with woodpecker
holes are preferred . In Louisiana eastern screech-owls chose nest
cavities in trees with lianas more often than expected according to
availability . Mean minimum diameter of cavity trees is 12 inches
(30 cm) . Eastern screech-owls usually choose cavities with
openings 3 to 5 inches (7.6-12.7 cm) in diameter. Typically, cavity
height is between 5 and 30 feet (1.5-9 m) above ground . Nest
cavities are usually reused in successive years . Nest boxes are
often reused by the same pair; one female was recorded using the same
box 8 years in a row .
Roosting: Eastern screech-owls prefer to roost in natural hollows
protected from rain and snow, and out of direct sunlight. They avoid
hollows used by fox squirrels (Sciurus niger) [7,19]. Roosting hollows
are typically 15 to 20 feet (4.5-6 m) and usually less than 40 feet (12
m) above the ground. Roosting cavity openings are about the same size
as nest cavity openings .
FOOD HABITS :
The eastern screech-owl is an opportunistic feeder. Prey species are
primarily small rodents, but numerous other animals are consumed .
Norberg  stated that insects are the major portion of the diet of
Megascops owls, but since the eastern screech-owl is sedentary it switches to
mammalian and avian prey in winter . The mammalian prey list
includes mice and rats (Muridae), shrews (Soricidae), moles (Talpidae),
flying squirrels (Glaucomys spp.), chipmunks (Tamias spp.), and bats
(Chiroptera). Avian prey includes songbirds, rock dove (Columbia
livia), northern bobwhite (Colinus virginianus), ruffed grouse (Bonasa
umbellus), American woodcock (Scolopax minor), common snipe (Gallinago
gallinago), American kestrel (Falco sparverius), and even other eastern
screech-owls . Eastern screech-owls have killed domestic pigeons,
hens, and ducks with prandial intent . Other vertebrate prey
includes snakes, lizards, frogs, toads, salamanders, and small fish.
Invertebrate prey includes many insects, snails, crayfish, spiders,
scorpions, millipedes, and earthworms .
Even though the prey lists are long, a few species form the main diet.
Favored prey includes meadow vole (Microtus pennsylvanicus),
white-footed mouse (Peromyscus leucopus), house mouse (Mus musculus),
and woodrats (Neotoma spp.) . Snyder and Wiley  estimated that
year-round prey consumption by eastern screech-owls is 30.7 percent
invertebrates, 0.6 percent lower vertebrates, 65.6 percent mammals, and
3.3 percent birds.
Seasonal Diet: In Michigan pellet analysis during spring and summer
indicated a high proportion of meadow vole, white-footed mouse,
crayfish, and small birds in eastern screech-owl diets . Pellet
analysis must be interpreted with caution, since many invertebrate prey
items do not contribute to pellet formation. In Ohio almost 65 percent
of the food items brought to nest boxes were birds, the majority of
which were migratory songbirds .
Craighead and Craighead's  analysis of pellets (Michigan study area)
indicated that meadow vole formed the bulk of the eastern screech-owl
winter diet. In a year when meadow vole numbers were lower than
average, a higher percentage of small birds appeared in pellets. In
both years, white-footed mouse were consumed in amounts comparable to
those of meadow voles. Other major pellet components were shrews and
moles . In Ohio fall and winter diets contained 30 percent birds
Foraging Habits: Eastern screech-owls tend to follow approximately the
same hunting route within the territory, tending to visit previously
successful sites . They usually forage in a perch-and-pounce manner,
taking short flights to capture already-spotted prey. Eastern
screech-owls have been observed to take a position near a bat colony and
capture bats as they leave the colony . Insects are often caught in
flight. Eastern screech-owls also walk on the ground to forage for
insects, particularly night-dormant grasshoppers, and wade into shallow
water after fish and aquatic invertebrates [18,21,37,39].
Larger owls prey on eastern screech-owl; the primary predator is the
great horned owl (Bubo virginianus), but others include barred owl
(Strix varia), great gray owl (S. nebulosa), snowy owl (Nyctea
scandiaca), long-eared owl (Asio otus), short-eared owl (A. flammeus),
and common barn owl (Tyto alba). Large hawks also take eastern
screech-owl, but only rarely since hawks are largely diurnal. Other
predators include domestic cat (Felis catus), Mustelids (minks, weasels,
and skunks), northern river otter (Lutra canadensis), bobcat (Lynx
rufus), and domestic dogs (Canis familiaris). Snakes have killed adult
eastern screech-owls (usually because an eastern screech-owl attacked a
snake too large to capture easily) but are more commonly nest predators
MANAGEMENT CONSIDERATIONS :
Population Status: Census of eastern screech-owl by song playback
provides a quick and relatively easy method to obtain accurate numbers
. Eastern screech-owl is common in the eastern United States, where
it is often the most common raptor [10,29]. Eastern screech-owl
population densities vary with the relative proportion of woodland cover
in an area . In Connecticut long-term survey data indicated a
decline in eastern screech-owl populations in the 1970's and 1980's,
possibly as a result of increased habitat fragmentation and conversion
of open woods to residential areas . In Ohio the eastern
screech-owl population fluctuated during the 30 years prior to 1975, but
no long-term trend was apparent in owls studied in suitable habitat.
The estimated annual recruitment needed to maintain population size is
2.21 fledglings per nesting pair. Eastern screech-owls have probably
increased with the opening up of dense forest but may be threatened by
increasing conversion of woods to residential areas . The increase
in early successional forest area in the latter half of the twentieth
century, largely due to abandonment of agricultural lands, may benefit
eastern screech-owls. Management considerations for maintenance of
populations include provision of natural and artificial cavities and
preservation of woodlots in suburban areas . Recommended size for
eastern screech-owl nest boxes: floor 8 inches by 8 inches (20 cm X 20
cm), cavity depth 8 to 10 inches (20-24 cm), and an entrance hole 3
inches (7.5 cm) in diameter .
Pesticide Accumulation: In the early 1970's eggshell thinning and DDE
and PCB levels in eggs were found to be relatively low for eastern
screech-owls. It was suggested that the consumption of rodents (lower
trophic levels) prior to egg-laying contributed to the relatively low
pesticide loads in eggs. Consumption of birds (higher trophic levels)
increased after eggs were incubated, and thus did not contribute to egg
pesticide loads . Other pesticides have not been investigated in
relationship to eastern screech-owl.
Predator-Prey Relationships: As a general rule, predators that are
specialized on small rodents tend to destabilize rodent population
cycles, whereas generalized predators tend to have a stabilizing effect,
sometimes suppressing cycles altogether. The eastern screech-owl is
more generalist in its effects on rodent populations . Even though
eastern screech-owls sometimes prey heavily on songbirds, they are more
likely to help keep rodent and insect populations in check .
Orchard owners reported that putting up nest boxes for screech-owls
resulted in a decrease in rodent damage to orchard trees . Eastern
screech-owls commonly nest in boxes put up for wood ducks (Aix sponsa),
American kestrels, squirrels, and purple martins (Progne subis) and will
even attempt to nest in mailboxes. Wood ducks often use nest boxes
after eastern screech-owls have left the nest; VanCamp and Henny 
suggested that wood duck nesting success is enhanced by eastern
screech-owl presence because owls discourage the usurpation of nest
boxes by European starlings (Sternus vulgaris).
Mortality Rates and Causes of Mortality: Mortality rate for the eastern
screech-owl was estimated as 69.5 percent the first year; eastern
screech-owls over 1 year old experience approximately 34 percent annual
mortality . Predation accounts for substantial adult mortality
. Collision with motor vehicles is also a major cause of mortality.
Other causes include shooting (which has become less frequent since
legal protection of raptors was enacted), electrocution, collision with
windows, or entrapment in buildings, and drowning (apparently these owls
take many baths and can become waterlogged and drown in rain barrels)
. There is a small amount of sibling cannibalism, usually after
accidental injury to a nestling . In Michigan nest loss due to fox
squirrels (Sciurus niger), which usually then appropriate the nest
hollow, has been observed .
FIRE EFFECTS AND USE
WILDLIFE SPECIES: Megascops asio
DIRECT FIRE EFFECTS ON ANIMALS :
There is no explicit information in the literature on the direct effect
of fire on eastern screech-owls. Adult birds are likely to escape fire.
Nestlings and eggs may be vulnerable.
HABITAT RELATED FIRE EFFECTS :
Habitat related fire effects include loss of habitat, change in habitat
structure (which may affect both nesting and hunting), and effects on
the prey base. In the southeastern states a majority of small mammals
thrive in early- to mid-successional habitats which are maintained by
fire or other disturbance . Prey vulnerability depends at least
partly on vegetation structure. Low, open vegetation makes prey easier
to spot, but eastern screech-owls require perches from which to hunt.
Fire-caused wounds may increase decay in large trees, making them more
likely to develop cavities (either natural or excavated).
FIRE USE :
FIRE REGIMES :
Find fire regime information for the plant communities in which this
species may occur by entering the species name in the FEIS home page under
"Find Fire Regimes".
WILDLIFE SPECIES: Megascops asio
1. Adam, Christopher I. G. 1987. Status of the eastern screech owl in
Saskatchewan with reference to adjacent areas. In: Nero, Robert W.;
Clark, Richard J.; Knapton, Richard J.; Hamre, R. H., eds. Biology and
conservation of northern forest owls: symposium proceedings; 1987
February 3-7; Winnipeg, MB. Gen. Tech. Rep. RM-142. Fort Collins, CO:
U.S. Department of Agriculture, Forest Service, Rocky Mountain Forest
and Range Experiment Station: 268-276. 
2. American Ornithologists' Union. 1957. Checklist of North American birds.
5th ed. Baltimore, MD: The Lord Baltimore Press, Inc. 691 p. 
3. American Ornithologists' Union. 1983. Checklist of North American birds.
6th ed. Lawrence, KS: Allen Press, Inc. 877 p. 
4. Belthoff, James R.; Ritchison, Gary. 1990. Roosting behavior of
postfledging eastern screech-owls. Auk. 107(3): 567-579. 
5. Bent, Arthur Cleveland. 1961. Life histories of North American birds of
prey. Part 2. New York: Dover Publications, Inc. 482 p. 
6. Call, Mayo. 1979. Habitat management guides for birds of prey. Techical
Note 338. Denver, CO: U.S. Department of the Interior, Bureau of Land
Management, Denver Service Center. 70 p. 
7. Craighead, John J.; Craighead, Frank C., Jr. 1969. Hawks, owls and
wildlife. Dover Edition. New York: Dover Publications, Inc. 443 p.
8. Cringan, Alexander T.; Horak, Gerald C. 1989. Effects of urbanization on
raptors in the western United States. In: Proceedings of the western
raptor management symposium and workshop; 1987 October 26-28; Boise, ID.
Scientific and Technical Series No. 12. Washington, DC: National
Wildlife Federation: 219-288. 
9. de la Torre, Julio. 1990. Owls: Their life and behavior. New York: Crown
Publishers. 214 p. 
10. DeGraaf, Richard M.; Scott, Virgil E.; Hamre, R. H.; [and others]. 1991.
Forest and rangeland birds of the United States: Natural history and
habitat use. Agric. Handb. 688. Washington, DC: U.S. Department of
Agriculture, Forest Service. 625 p. 
11. DeGraaf, Richard M.; Yamasaki, Mariko; Leak, William B.; Lanier, John W.
1992. New England wildlife: management of forested habitats. Gen. Tech.
Rep. NE-144. Radnor, PA: U.S. Department of Agriculture, Forest Service,
Northeastern Forest Experiment Station. 271 p. 
12. Emmerich, John M.; Vohs, Paul A. 1982. Comparative use of four woodland
habitats by birds. Journal of Wildlife Management. 46(1): 43-49.
13. Fitton, Sam. 1993. Screech-owl distribution in Wyoming. Western Birds.
24: 182-188. 
14. Gehlbach, Frederick R. 1986. Odd couples of suburbia. Natural History.
95(6): 56-64, 66. 
15. Hamel, Paul B.; LeGrand, Harry E., Jr.; Lennartz, Michael R.;
Gauthreaux, Sidney A., Jr. 1982. Bird-habitat relationships on
southeastern forest lands. Gen. Tech. Rep. SE-22. Asheville, NC: U.S.
Department of Agriculture, Forest Service, Southeastern Forest
Experiment Station. 417 p. 
16. Hardin, Kimberly I.; Evans, Keith E. 1977. Cavity nesting bird habitat
in the oak-hickory forests--a review. Gen. Tech. Rep. NC-30. St. Paul,
MN: U.S. Department of Agriculture, Forest Service, North Central Forest
Experiment Station. 23 p. 
17. Harlow, Richard F.; Guynn, David C., Jr. 1983. Snag densities in managed
stands of the South Carolina coastal plain. Southern Journal of Applied
Forestry. 7(4): 224-229. 
18. Johnsgard, Paul A. 1988. North American owls. Washington, DC:
Smithsonian Institute Press. 295 p. 
19. Karalus, Karl E.; Eckert, Allan W. 1974. The owls of North America
(north of Mexico). Garden City, NY: Doubleday & Company, Inc. 278 p.
20. Kochert, Michael N. 1986. Raptors. In: Cooperrider, Allan Y.; Boyd,
Raymond J.; Stuart, Hanson R., eds. Inventory and monitoring of wildlife
habitat. Denver, CO: U.S. Department of the Interior, Bureau of Land
Management, Denver Service Center: 313-349. 
21. Marshall, Joe T., Jr. 1967. Parallel variation in North and Middle
American screech-owls. Monongraphs of the Western Foundation of
Vertebrate Zoology No. 1. Los Angles, CA: Western Foundation of
Vertebrate Zoology. 72 p. 
22. Marti, Carl D.; Marks, Jeffrey S. 1989. Medium-sized owls. In:
Proceedings of the western raptor management symposium and workshop;
1987 October 26-28; Boise, ID. Scientific and Technical Series No. 12.
Washington, DC: National Wildlife Federation: 124-133. 
23. Mazur, Kurt M. 1992. Fall food of the eastern screech-owl in Manitoba.
Blue Jay. 50(1): 33-35. 
24. McComb, William C.; Noble, Robert E. 1981. Nest-box and natural-cavity
use in three mid-south forest habitats. Journal of Wildlife Management.
45(1): 93-101. 
25. Norberg, R. Ake. 1987. Evolution, structure, and ecology of northern
forest owls. In: Nero, Robert W.; Clark, Richard J.; Knapton, Richard
J.; Hamre, R. H., eds. Biology and conservation of northern forest owls:
symposium proceedings; 1987 February 3-7; Winnipeg, MB. Gen. Tech. Rep.
RM-142. Fort Collins, CO: U.S. Department of Agriculture, Forest
Service, Rocky Mountain Forest and Range Experiment Station: 9-43.
26. Schroeder, Richard L.; Cable, Ted T.; Haire, Sandra L. 1992. Wildlife
species richness in shelterbelts: test of a habitat model. Wildlife
Society Bulletin. 20(3): 264-273. 
27. Sibley, Charles G.; Monroe, Burt L., Jr. 1990. Distribution and taxonomy
of the birds of the world. New Haven, CT: Yale University Press. 1111 p.
28. Smith, Dwight G. 1993. Eastern screech owls. Bird Watcher's Digest.
16(1): 32-39. 
29. Smith, Dwight G.; Devine, Arnold; Walsh, Dan. 1987. Censusing screech
owls in southern Connecticut. In: Nero, Robert W.; Clark, Richard J.;
Knapton, Richard J.; Hamre, R. H., eds. Biology and conservation of
northern forest owls: symposium proceedings; 1987 February 3-7;
Winnipeg, MB. Gen. Tech. Rep. RM-142. Fort Collins, CO: U.S. Department
of Agriculture, Forest Service, Rocky Mountain Forest and Range
Experiment Station: 255-267. 
30. Smith, Dwight G.; Gilbert, Raymond. 1984. Eastern screech-owl home range
and use of suburban habitats in southern Connecticut. Journal of Field
Ornithology. 55(5): 322-329. 
31. Snyder, Noel F. R.; Wiley, James W. 1976. Sexual size dimorphism in
hawks and owls of North America. Ornithological Monographs. No. 20.
Gainesville, FL: American Ornithologists' Union. 96 p. 
32. Sparks, Earl J.; Belthoff, James R.; Ritchison, Gary. 1994. Habitat use
by eastern screech-owls in central Kentucky. Journal of Field
Ornithology. 65(1): 83-95. 
33. Stewart, Robert E.; Robbins, Chandler S. 1958. Birds of Maryland and the
District of Columbia. North American Fauna: No. 62. Washington, DC: U.S.
Department of the Interior, Fish and Wildlife Service. 401 p. 
34. Tate, James, Jr. 1986. The Blue List for 1986. American Birds. 40(2):
35. Terres, John K. 1980. The Audubon Society encyclopedia of North American
birds. New York: Alfred A. Knopf. 1109 p. 
36. Tyler, Hamilton, A.; Phillips, Don. 1978. Owls by day and night. Happy
Camp, CA: Naturegraph Publishers, Inc. 208 p. 
37. VanCamp, Laurel F.; Henny, Charles J. 1975. The screech owl: its life
history and population ecology in northern Ohio. North American Fauna,
Number 71. Washington, DC: U.S. Department of the Interior, Fish and
Wildlife Service. 65 p. 
38. Lynch, Patrick J.; Smith, Dwight G. 1984. Census of eastern screech-owls
(Otus asio) in urban open-space areas using tape-recorded song. American
Birds. 38: 388-391. 
39. Allen, A. A. 1924. A contribution to the life history and economic
status of the screech owl (Otus asio). Auk. 41(1): 1-17. 
40. Landers, J. Larry. 1987. Prescribed burning for managing wildlife in
southeastern pine forests. In: Dickson, James G.; Maughan, O. Eugene,
eds. Managing southern forests for wildlife and fish: a proceedings;
[Date of conference unknown]; [Location of conference unknown]. Gen.
Tech. Rep. SO-65. New Orleans, LA: U.S. Department of Agriculture,
Forest Service, Southern Forest Experiment Station: 19-27. 
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