Richard T. Reynolds
Research Wildlife Biologist
240 West Prospect Road
Fort Collins, CO 80526
1. Determine the distribution and density of breeding territories, reproduction, survival, mate and territory fidelity, recruitment, and emigration/immigration of northern goshawks on the Kaibab Plateau, Arizona. 2. Assess the relationship between goshawk demographic performance and habitat conditions on territories. 3. Identify the compositional and sturctural habitat elements that confer quality to goshawk breeding habitats. 4. Investigate post-fledgling movements and dispersal of juvenile goshawks.5. Determine the causes and consequences of dispersal of adult, breeding goshawks. 6. Identify factors affecting goshawk prey populations and the extent to which prey affects goshawk demographics. 7. Investigate the utility of genetic fingerprinting for identifying individual goshawks from in capture-recapture studies.8. Investigate the utility of using color, pattern, and shape of molted feathers for for identifying individual goshawks in mark-recapture studies. 9. Determine the effects of high vs low intensity fire on the habitat on goshawk prey populations and on goshawk habitat use.
My research focuses on the relationship between species' demographies -- their reproduction, survival, mate and territory fidelity -- and their habitat in order to identify habitat elements that distinguish among high quality and low quality habitats. This research necessarily requires long-term capture-mark-recapture work so that the demographic performance of known individuals can be related to the compositions and structures of habitats they use throughout their lives. Products of the research enhance our understanding of the importance of habitat characteristics in species' viability and helps managers provide quality habitats through management activities.
An example of this demographically-based habitat research is our work on the northern goshawk, a 'sensitive' species and a candidate for listing under the Endangered Species Act. The focus of concern for this species was loss of habitat due to a forest management practices. The importance of our research on the hawk and its habitat was highlighted in a collaborative effort between animal ecologists, silviculturalists, and foresters to synthesize habitat-related knowledge of the plants and animals in this predator's food web to develop best-science management recommendations. This collaboration identified unique sets of desired habitat conditions that combined both a fine and coarse filter approach to multi-species conservation. The management recommendations were incorporated into forest plans of all National Forests in the Southwestern Region in 1996. Implementation improves visual quality, forage, wood products, and hydrologic function of forests; reduces fire hazards; and increases resilience to insects, disease, and climate change.
Why This Research is Important
1. Initiated, conducted, and continue to collaborate on a 31-year capture-recapture investigation of the relationship between the habitat composition and structure and the demography of the flammulated owl in Colorado ponderosa pine forests.2. Initiated and conducted a 10-year study of the distribution, density, territory occupancy, and reproduction of Mexican spotted owls in Colorado.3. Initiated and conducted a 4-year study of birds , mammals, and herps in aspen and spruce-fir forests in Colorado and Wyoming. 4. Collobratively initiated and conducted a northern goshawk habitat assessment for the entire Black Hills National Forest.
- Oregon State University, BS Botony, Entomology, Zoology, History and Philosophy of Science, 1970
- Oregon State University, MS Wildlife Ecology, 1975
- Oregon State University, Ph.D. Animal Ecology, 1978
Featured Publications & Products
- Reynolds, Richard T.; Graham, Russell T.; Boyce, Douglas A., Jr. 2008. Northern goshawk habitat: an intersection of science, management, and conservation.
- Youtz, James A.; Graham, Russell T.; Reynolds, Richard T.; Simon, Jerry. 2008. Implementing northern goshawk habitat management in Southwestern forests: a template for restoring fire-adapted forest ecosystems..
- Bayard De Volo, Shelley; Reynolds, Richard T.; Douglas, Marlis R.; Antolin, Michael F. 2008. An improved extraction method to increase DNA yield from molted feathers.
- Linkhart, Brian D.; Reynolds, Richard T. 2007. Return rate, fidelity, and dispersal in a breeding population of flammulated owls (Otus flammeolus).
- Salafsky, Susan R.; Reynolds, Richard T.; Noon, Barry R.; Wiens, John A. 2007. Reproductive responses of northern goshawks to variable prey populations.
Publications & Products
- Dickson, Brett G.; Sisk, Thomas D.; Sesnie, Steven E.; Reynolds, Richard T.; Rosenstock, Steven S.; Vojta, Christina D.; Ingraldi, Michael F.; Rundall, Jill M. 2014. Integrating single-species management and landscape conservation using regional habitat occurrence models: The northern goshawk in the Southwest, USA.
- Bayard de Volo, Shelley; Reynolds, Richard T.; Sonsthagen, Sarah A.; Talbot, Sandra L.; Antolin, Michael F. 2013. Phylogeography, postglacial gene flow, and population history of North American northern goshawks (Accipiter gentilis).
- Reynolds, Richard T.; Sanchez Meador, Andrew J.; Youtz, James A.; Nicolet, Tessa; Matonis, Megan S.; Jackson, Patrick L.; DeLorenzo, Donald G.; Graves, Andrew D. 2013. Restoring composition and structure in Southwestern frequent-fire forests: A science-based framework for improving ecosystem resiliency.
- Reynolds, Richard T.; Boyce, Douglas A.; Graham, Russell T. 2012. Ponderosa pine forest structure and northern goshawk reproduction: Response to Beier et al.
- Boyce, Douglas A., Jr..; Reynolds, Richard T.; Graham, Russell T. 2006. Goshawk status and management: What do we know, what have we done, where are we going?.
- Wiens, J. David; Reynolds, Richard T.; Noon, Barry R. 2006. Juvenile movement and natal dispersal on northern goshawks in Arizona.
- Wiens, J. David; Noon, Barry R.; Reynolds, Richard T. 2006. Post-fledging survival of Northern Goshawks: the importance of prey abundance, weather, and dispersal.
- Wiens, J. David; Reynolds, Richard T. 2005. Is fledging success a reliable index of fitness in Northern Goshawks?.
- Salafsky, Susan R.; Reynolds, Richard T.; Noon, Barry R. 2005. Patterns of temporal variation in goshawk reproduction and prey resources.
- de Volo, Shelley Bayard; Reynolds, Richard T.; Topinka, J. Rick; May, Bernie; Antolin, Michael F. 2005. Population genetics and genotyping for mark-recapture studies of Northern Goshawks (Accipiter gentilis) on the Kaibab Plateau, Arizona.
- Reynolds, Richard T.; Wiens, J. David; Joy, Suzanne M.; Salafsky, Susan R. 2005. Sampling considerations for demographic and habitat studies of Northern Goshawks.
- La Sorte, Frank A.; Mannan, R. William; Reynolds, Richard T.; Grubb, Teryl G. 2004. Habitat associations of sympatric red-tailed hawks and northern goshawks on the Kaibab Plateau.
- Joy, Suzanne M.; Reich, R. M.; Reynolds, Richard T. 2003. A non-parametric, supervised classification of vegetation types on the Kaibab National Forest using decision trees.
- Johnson, Charles L.; Reynolds, Richard T. 2002. Responses of Mexican spotted owls to low-flying military jet aircraft.
- Belnap, Jayne; Reynolds, Richard; Reheis, Marith; Phillips, Susan L. 2001. What makes the desert bloom? Contribution of dust and crusts to soil fertility on the Colorado Plateau.
- Joy, Suzanne M.; Reich, Robin M.; Reynolds, Richard T. 2000. Modeling small-scale variability in the composition of goshawk habitat on the Kaibab National Forest.
- Reynolds, Richard T.; Joy, Suzanne M. 1998. Distribution, territory occupancy, dispersal, and demography of northern goshawks on the Kaibab Plateau, Arizona.
- Linkhart, Brian D.; Reynolds, Richard T. 1997. Territories of flammulated owls (Otus flammeolus): Is occupancy a measure of habitat quality?.
- Joy, Suzanne M.; Reynolds, Richard T.; Knight, Richard L.; Hoffman, Richard W. 1994. Feeding ecology of sharp-shinned hawks in deciduous and coniferous forests in Colorado.
- Joy, Suzanne M.; Reynolds, Richard T.; Leslie, Douglas G. 1994. Northern goshawk broadcast surveys: Hawk response variables and survey cost.
- Reynolds, Richard T.; Graham, Russell T.; Reiser, M. Hildegard. 1992. Management recommendations for the northern goshawk in the southwestern United States.
- Reynolds, Richard T.; Linkhart, Brian D. 1990. Longevity records for male and female flammulated owls.
- Linkhart, Brian D.; Reynolds, Richard T. 1987. Brood division and postnesting behavior of Flammulated Owls.
- Reynolds, Richard T.; Linkhart, Brian D. 1984. Methods and materials for capturing and monitoring flammulated owls.
- Reynolds, Richard T. 1983. Management of western coniferous forest habitat for nesting accipiter hawks.
- Reynolds, Richard T.; Wight, Howard M. 1978. Distribution, density, and productivity of accipiter hawks breeding in Oregon.
- Reynolds, Richard T. 1970. Nest observations of the long-eared owl (Asio otus) in Benton County, Oregon, with notes on their food habits.