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Introduction
Understanding the natural regeneration process for whitebark pine (Pinus albicaulis) is essential for managing whitebark pine communities.
It provides insight into the successional dynamics of whitebark pine, how succession varies geographically, and particularly how fire may be used as a management tool to return whitebark pine communities to earlier successional stages.
Knowledge of the regeneration process is integral to all management strategies for whitebark pine including approaches to managing for white pine blister rust (Cronartium ribicola), whether for enhancing natural resistance or for growing rust-resistant seedlings.
Natural Regeneration
The natural regeneration process of whitebark pine may take up to five years to complete. There are three distinct phases of the natural regeneration process; 1) cone and seed initiation, development, and maturation; 2) seed dissemination; and 3) germination.
The over two-year period for cone initiation, development and maturation of whitebark pine is similar to that of most pine species worldwide, including the European and Asian stone pines of the Pinus subsection Cembrae.
This process begins in whitebark pine trees between the ages of 20 and 30 years although large crops don't occur until at least 60 to 80 years of age when trees have wide extensive crowns.
Developmental Stages
Several developmental stages of the cones and seeds occur during the first phase, starting with the initiation of staminate and ovulate cone buds and ending with mature cones and seeds.
An important second phase is the dispersal and burial of seeds, commonly referred to as "caching", by animals, primarily Clark's nutcrackers (Nucifraga columbiana), which is described in another chapter but is briefly summarized here.
The final phase of the natural regeneration process includes the physiological development leading up to seed germination, the emergence process, and establishment of young seedlings.
Related Links
Photographs
Click on each thumbnail photograph for an enlarged version.
![[Photograph]: Typical multiple stem growth-form with a dense spreading crown shape.](/rm/ecology/studies/whitebark/images/image002.jpg) Typical multiple stem growth-form with a dense spreading crown shape. |
![[Photograph]: Seeds of whitebark pine are important for several wildlife species including grizzly bear.](/rm/ecology/studies/whitebark/images/image004.jpg) Seeds of whitebark pine are important for several wildlife species including grizzly bear. |
![[Photograph]: The Clarks Nutcracker is the primary seed dispersal agent for whitebark pine.](/rm/ecology/studies/whitebark/images/image006.jpg) The Clarks Nutcracker is the primary seed dispersal agent for whitebark pine. |
![[Photograph]: Whitebark pine seedling clusters range from one to more than 20 per cluster.](/rm/ecology/studies/whitebark/images/image008.jpg) Whitebark pine seedling clusters range from one to more than 20 per cluster. |
![[Photograph]: Planted whitebark seedlings often have multiple tops.](/rm/ecology/studies/whitebark/images/image010.jpg) Planted whitebark seedlings often have multiple tops. |
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