Alejandro A. Royo
PO Box 267
Irvine, PA 16329
My main research focus investigates the role of competition by understory plants and herbivory by ungulates and small mammals in the diversity and abundance of tree seedlings in the understory before overstory gap formation. Furthermore, I am involved in several projects elucidating the role mammalian herbivores play in the maintenance of herbaceous plant communities across a wide latitudinal gradient.
I plan on continuing my research programs on the maintenance of herbaceous species diversity in both temperate and tropical systems. I also plan on examining the potential for developing alternatives to herbicide application for the control of understory herbaceous interference. Additional research interests involve regeneration of forest systems following catastrophic wind disturbance and subsequent salvaging as well as the impact of soil-borne pathogens on early tree establishment.
Why This Research is Important
Among the many factors that determine forest understory diversity prior to any overstory disturbance, competition and herbivory are typically thought of as preeminent. Understanding how these two factors, alone and in concert, act to promote or reduce plant diversity is key to the sustainable management of forest resources.
- University of Pittsburgh, Ph.D. Ecology and Evolutionary Biology, 2005
- University of North Carolina, Greensboro, M.S. Department of Biological Sciences, 1998
- University of North Carolina, Greensboro, B.A. Department of Biological Sciences, 1993
- University of Pittsburgh, Graduate Certificate Latin American Studies, 2005
- Ecological Society of America
Featured Publications & Products
- Royo, Alejandro A.; Heartsill Scalley, Tamara; Moya, Samuel; Scatena, Fred N. 2011. Non-arborescent vegetation trajectories following repeated hurricane disturbance: ephemeral versus enduring responses.
- Royo, Alejandro A.; Carson, Walter P. 2010. The formation of dense understory layers in forests worldwide: consequences and implications for forest dynamics, biodiversity, and succession.
- Royo, Alejandro A.; Stout, Susan L.; deCalesta, David S.; Pierson, Timothy G. 2010. Restoring forest herb communities through landscape-level deer herd reductions: Is recovery limited by legacy effects?.
- Royo, Alejandro A.; Collins, Rachel; Adams, Mary Beth; Kirschbaum, Chad; Carson, Walter P. 2010. Pervasive interactions between ungulate browsers and disturbance regimes promote temperate forest herbaceous diversity.
- Knight, Kathleen S.; Herms, Daniel A.; Cardina, John; Long, Robert; Rebbeck, Joanne; Gandhi, Kamal J.K.; Smith, Annemarie; Klooster, Wendy S.; Herms, Catherine P.; Royo, Alejandro A. 2010. Emerald ash borer aftermath forests: The dynamics of ash mortality and the responses of other plant species.
Publications & Products
- Huebner, Cynthia D.; Steinman, Jim; Hutchinson, Todd F.; Ristau, Todd E.; Royo, Alejandro A. 2014. The distribution of a non-native (Rosa multiflora) and native (Kalmia latifolia) shrub in mature closed-canopy forests across soil fertility gradients.
- Peterson, Chris J.; Krueger, Lisa M.; Royo, Alejandro A.; Stark, Scott; Carson, Walter P. 2013. Disturbance size and severity covary in small and mid-size wind disturbances in Pennsylvania northern hardwoods forests.
- Nuttle, Tim; Royo, Alejandro A.; Adams, Mary Beth; Carson, Walter P. 2013. Historic disturbance regimes promote tree diversity only under low browsing regimes in eastern deciduous forest.
- Royo, Alejandro A.; Ristau, Todd E. 2013. Stochastic and deterministic processes regulate spatio-temporal variation in seed bank diversity.
- Stout, Susan L.; Royo, Alejandro A.; deCalesta, David S.; McAleese, Kevin; Finley, James C. 2013. The Kinzua Quality Deer Cooperative: can adaptive management and local stakeholder engagement sustain reduced impact of ungulate browsers in forest systems?.
- Huebner, Cynthia D.; Hutchinson, Todd; Ristau, Todd; Royo, Alejandro; Steinman, James. 2012. Documenting the Regional and local distribution of Kalmia latifolia and Rosa multiflora in West Virginia, Ohio, and Pennsylvania Forests along a soil fertility gradient.
- Dey, Daniel C.; Royo, Alejandro A.; Gardiner, Emile S.; Cardenas, Luis Mario. 2012. Sustaining Quercus humboltii and Colombobalanus excelsa on the Colombian landscape: preservation or conservation - a research perspective.
- Dey, Daniel C.; Royo, Alejandro A.; Gardiner, Emile S.; Cardenas, Luis Mario. 2012. Sustaining Quercus humboldtii and Colombobalanus excelsa on the Colombian landscape: preservation or conservation - a research perspective.
- Royo, Alejandro A.; Knight, Kathleen S. 2012. White ash (Fraxinus americana) decline and mortality: the role of site nutrition and stress history.
- Royo, Alejandro A.; Knight, Kathleen S.; Himes, Jamie M.; Will, Ashley N. 2012. White ash (Fraxinus americana) health in the Allegheny plateau region, Pennsylvania: Evaluating the relationship between FIA phase 3 crown variables and a categorical rating system.
- Goetsch, Chandra; Wigg, Jennifer; Royo, Alejandro A.; Ristau, Todd; Carson, Walter P. 2011. Chronic over browsing and biodiversity collapse in a forest understory in Pennsylvania: Results from a 60 year-old deer exclusion plot.
- Kain, Morgan; Battaglia, Loretta; Royo, Alejandro; Carson, Walter P. 2011. Over-browsing in Pennsylvania creates a depauperate forest dominated by an understory tree: results from a 60-year-old deer exclosure.
- Dey, Daniel C.; Royo, Alejandro A.; Brose, Patrick H.; Hutchinson, Todd F.; Spetich, Martin A.; Stoleson, Scott H. 2010. An ecologically based approach to oak silviculture: a synthesis of 50 years of oak ecosystem research in North America.
- Reinhart, Kurt O.; Royo, Alejandro A.; Kageyama, Stacie A.; Clay, Keith. 2010. Canopy gaps decrease microbial densities and disease risk for a shade-intolerant tree species.
- Krueger, Lisa M.; Peterson, Chris J.; Royo, Alejandro; Carson, Walter P. 2009. Evaluating relationships among tree growth rate, shade tolerance, and browse tolerance following disturbance in an eastern deciduous forest.
- Royo, Alejandro A.; Bates, Ramona M.; Lacey, Elizabeth P. 2008. Demographic constraints in three populations of Lobelia boykinii, a rare wetland endemic.
- Royo, Alejandro A.; Carson, Walter P. 2008. Direct and indirect effects of a dense understory on tree seedling recruitment in temperate forests: habitat-mediated predation versus competition.
- Royo, A.A. 2008. P.A. Thomas and J. Packham: Ecology of woodlands and forests: Description, dynamics, and diversity [book review].
- Royo, Alejandro A.; Carson, Walter P. 2006. On the formation of dense understory layers in forests worldwide: consequences and implications for forest dynamics, biodiversity, and succession.
- Comisky, Lauren; Royo, Alejandro A.; Carson, Walter P. 2005. Deer browsing creates rock refugia gardens on large boulders in the Allegheny National Forest, Pennsylvania.
- Banta, Joshua A.; Royo, Alejandro A.; Kirschbaum, Chad; Carson, Walter P. 2005. Plant communities growing on boulders in the Allegheny National Forest: Evidence for boulders as refugia from deer and as a bioassay of overbrowsing.
- Reinhart, Kurt O.; Royo, Alejandro; Van der Putten, Wim H.; Clay, Keith. 2005. Soil feedback and pathogen activity in Prunus serotina throughout its native range.
- Royo, Alejandro A.; Carson, Walter P. 2005. The herb community of a tropical forest in central Panama: dynamics and impact of mammalian herbivores.
|Can We Bank on Forest Seed Banks|
Community composition of seeds stored in forest soils becomes increasingly divergent over time
|Hurricanes Disturb Non-tree Subtropical Wet Forest Species Composition |
Hurricane disturbance caused pronounced and persistent changes in the non-tree species composition of a subtropical wet forest. A unique long-te ...
|Landscape-level Deer Herd Reductions Restore Forest Understory Plant Communities but Not Species Diversity|
Since 2001, NRS scientist Alejandro Royo has tracked the response of herbaceous plant communities to deer herd reductions throughout the 70,000- ...