Pacific Southwest Research Station

Selected Publications

Ashton, D. T., and H. H. Welsh, Jr. 2012 (In review). A conservation strategy for the western pond turtle (Actinemys marmorata) in California. Calif. Dept. Fish and Game.

Bury, R. B., H. H. Welsh, Jr., D. J. Germano, and D. T. Ashton (eds). 2012 (In press). Western Pond Turtle; biology, sampling techniques, inventory and monitoring, conservation and management. Northwest Fauna 7 (monograph series).

Wheeler, C., H. H. Welsh, Jr., and L. M. Ollivier. 2012 (In press). Reproductive biology of the Del Norte salamander (Plethodon elongatus) in northwest California. Journal of Herpetology.

Welsh, H. H., Jr. 2011. Frogs, fish, and forestry: an integrated watershed network paradigm conserves biodiversity and ecological services. Diversity 3:503-530.

Ashton, D. T., J. B. Bettaso, and H. H. Welsh, Jr. 2011. Comparative ecology of western pond turtle (Actinemys marmorata) populations on the free-flowing South Fork and regulated Main Fork, Trinity River: demography, size and body condition comparisons, thermal ecology, and spatial dynamics. Report to the Trinity River Restoration Program. Weaverville, CA.

Fuller, T., K. L Pope, D. T. Ashton, and H. H. Welsh, Jr. 2011. Linking the distribution of an invasive amphibian (Rana catesbeiana) to habitat conditions in a managed river system in northern California. Restoration Ecology 19:204-213.

Welsh, H. H., Jr. and G. R. Hodgson. 2010. Spatial relationships in a dendritic network: the herpetofaunal metacommunity of the Mattole River catchment of northwest California. Ecography 34:49-66.

Welsh, H. H., Jr., W. H. Clark, E. Franco-Vizcaino, and J. H. Vadez-Villavicencio. 2010. Herpetofauna associated with palm oases across the Californian-Sonoran transition in northern Baja California, Mexico. The Southwestern Naturalist 55:581-585.

Garwood, J. M. and H. H. Welsh, Jr. 2010. Thamnophis atratus hydrophilus. Maximum elevation. Herpetological Review 41:503-504. Natural History Note.

Welsh, H. H., Jr., G. R. Hodgson, J. J. Duda, and J. M. Emlen. 2010. Faunal Assemblages and multi-scale habitat patterns in headwater tributaries of the south fork Trinity River– an unregulated river embedded within a multi-use landscape. Animal Conservation and Biodiversity 33:63-87.

Welsh, H. H., Jr., C. A. Wheeler, and A. J Lind. 2010. Spatial ecology of the aquatic garter snake, Thamnophis atratus, in a free-flowing stream environment. Copeia 2010:75-85.

Clayton, D. R., L. M. Ollivier, and Welsh, H. H., Jr. 2009. Survey protocol for the Siskiyou Mountains salamander (Plethodon stormi). Version 3.0. Chapter 6 in: Olson, D. H., D. R. Clayton, R. S. Nauman, and H. H. Welsh, Jr. Conservation of the Siskiyou Mountains salamander (Plethodon stormi). Northwest Fauna 6:43-61.

Olson, D. H., D. R. Clayton, E. C. Reilly, R. S. Nauman, B. Devlin, and H. H. Welsh, Jr. 2009. Site identification. Chapter 4 in: Olson, D. H., D. R. Clayton, R. S. Nauman, and H. H. Welsh, Jr. 2009. Conservation of the Siskiyou Mountains salamander (Plethodon stormi). Northwest Fauna 6:31-37.

Reilly, E. C. Olson, D. H., D. R. Clayton, , R. S. Nauman, B. Devlin, and H. H. Welsh, Jr. 2009. Spatial model of optimal habitat for the Siskiyou Mountains salamander (Plethodon stormi), north of the Siskiyou Crest. Northwest Fauna 6:23-25.

Olson, D. H., D. R. Clayton, E. C. Reilly, R. S. Nauman, B. Devlin, and H. H. Welsh, Jr. 2009. Conservation strategy for the Siskiyou Mountains salamander (Plethodon stormi), Version 1.0. Chapter 1 in: Olson, D. H., D. R. Clayton, R. S. Nauman, and H. H. Welsh, Jr. 2009. Conservation of the Siskiyou Mountains salamander (Plethodon stormi). Northwest Fauna 6:1-22.

Olson, D. H., D. R. Clayton, R. S. Nauman, and H. H. Welsh, Jr. (Editors). 2009. Conservation of the Siskiyou Mountains salamander (Plethodon stormi). Northwest Fauna 6. Olympia, WA; Society of Northwestern Vertebrate Biology.

Welsh, H. H., Jr. 2009. Northwestern California, pp.75-77, in: Olson. D. H. (coord. ed.). Herpetological conservation in northwestern North America. Northwestern Naturalist 90:61-96.

Welsh, H. H., Jr., and Garth R. Hodgson. 2009. Stream Amphibians as metrics of critical biological thresholds in the Pacific Northwest, U.S.A: a response to Kroll et al. Journal of Freshwater Biology.

Welsh, H. H., Jr., K. L. Pope and C.A. Wheeler. 2009. How reliable are amphibian population metrics: A response to Kroll et al. Biological Conservation.

Pope, K. L., J. M. Garwood, H. H. Welsh, Jr. and S. P. Lawler. 2008. Evidence of indirect impacts of introduced trout on native amphibians via facilitation of a shared predator. Biological Conservation, 141:1321-1331.

Wheeler, C. A., and H. H. Welsh, Jr. 2008. Mating strategy and breeding patterns of the foothill yellow-legged frog (Rana boylii). Herpetological Conservation and Biology 3(2):128-142.

Welsh, H. H., Jr., K. L. Pope and C.A. Wheeler. 2008. Using multiple metrics to assess the effects of forest succession on population status: A comparative study of two terrestrial salamanders in the US Pacific Northwest. Biological Conservation, 141: 1149-1160.

Wheeler, C. A, and H. H. Welsh, Jr. 2008. Mating strategy and breeding patterns of the foothill yellow-legged frog (Rana boylii). Herpetological Conservation and Biology, 3(2):128-142.

Welsh, H. H., Jr., and G. R. Hodgson. 2008. Amphibians as metrics of critical biological thresholds in forested headwater streams of the Pacific Northwest, USA. Journal of Freshwater Biology, 53: 1470-1488.

Fellers, G. M., K.L. Pope , J. E. Stead, M. S. Koo, and H.H. Welsh, Jr. 2008. Turning population trend monitoring into active conservation: Can we save the Cascades frog in the Lassen region of California. Herpetological Conservation and Biology 3:28-39.

Welsh, H. H., Jr., G. M. Fellers, and A. J. Lind. 2007. Amphibian populations in the terrestrial environment: is there evidence for declines of terrestrial forest amphibians in northwestern California. Journal of Herpetology 4:469-482.

Olson, D. H., P. D. Anderson, C. A. Frissell, H. H. Welsh, Jr. and D. F. Bradford. 2007. Biodiversity management approaches for stream riparian areas: perspectives for Pacific Northwest headwater forests and amphibians. Forest Ecology and Management 246:81-107.

Wheeler, C. A., N. E. Karraker, H. H. Welsh, Jr., and L. M. Ollivier. 2007. Diet of the Del Norte salamander (Plethodon elongatus): differences by age, gender, and season,. Northwestern Naturalist 88:85-94.

Welsh, H. H., Jr., H. Stauffer, D. Clayton, and L. Ollivier. 2007. Strategies for modeling habitat relationships of rare species: an example using the Siskiyou Mt. salamander (Plethodon stormi). Northwest Science 81:15-36.

Garwood, J., C. Wheeler, R. Bourque, M. Larson, and H. H. Welsh, Jr. 2007. Egg mass drift increases vulnerability during early development of Cascades frogs (Rana cascadae). Northwestern Naturalist 88:94-96.

Ashton, D. T., S. B. Marks, and H. H. Welsh, Jr. 2006. Evidence of continued effects from timber harvesting on lotic amphibians in redwood forests of northwestern California. Forest Ecology and Management 221: 183-193.

Welsh, H. H., Jr., J. R. Dunk, W. J. Zielinski. 2006. Developing and applying habitat models using forest inventory data: an example using a terrestrial salamander. Journal of Wildlife Management 70: 671-681.

Welsh, H. H., Jr., K. L. Pope, and D. Boiano. 2006. Sub-alpine amphibian distributions related to species palatability to non-native salmonids in the Klamath mountains of northern California. Diversity and Distributions 12: 298-309.

Dunk, J. R., W. J. Zielinski, and H. H. Welsh, Jr. 2006. Evaluating reserves for species richness and representation in northern California. Diversity and Distributions 12:434-442.

Karraker, N. E. and H. H. Welsh, Jr. 2006. Long-term impacts of even-aged timber management on abundance and body condition of terrestrial amphibians in Northwestern California. Biological Conservation 131: 132-140.

Spickler, J. C., S. C. Sillett, S. B. Marks and H. H. Welsh, Jr. 2006. Evidence of a new niche for a North American salamander: Aneides vagrans residing in the canopy of old-growth redwood forest. Herpetological Conservation and Biology 1: 16-27.

Welsh, H. H., Jr., G. R. Hodgson, and N. E. Karraker. 2005b. Influences of the vegetation mosaic on riparian and stream microclimates in a mixed forest-grassland landscape in "Mediterranean" northwestern California. Ecography 28: 537-551.

Welsh, H. H., Jr., G. R. Hodgson, and A. J. Lind. 2005a. Ecogeography of the herpetofauna of a northern California watershed: linking species patterns to landscape processes. Ecography 28: 521-536.

Bury, R. B., and H. H. Welsh, Jr. 2005. Siskiyou mountains salamander, Plethodon stormi. in M. Lanno, editor. Status and Conservation of U. S. Amphibians. U. C. Press, Berkeley, CA.

Feral, D., M. A. Camann, and H. H. Welsh, Jr. 2005. Dicamptodon tenebrosus larvae within hyporheic zones of intermittent streams in California. Herpetological Review 36:26-27.

Garwood, J. M., and H. H. Welsh, Jr. 2005. Rana cascadae (Cascades frog) predation. Herpetological Review 36:165.

Lind, A. J., H. H. Welsh, Jr., and D. A. Tallmon. 2005. Garter snake population dynamics from a 16-year study: considerations for ecological monitoring. Ecological Applications 15:294-303.

Wheeler, C. A., J. M. Garwood, and H. H. Welsh, Jr. 2005. Rana boylii (foothill yellow-legged frog) physiological skin color transformation. Herpetological Review 36:164-165.

Welsh, H. H., Jr. 2005. Pacific giant salamander, Dicamptodon tenebrosus. in M. Lanno, editor. Status and Conservation of U. S. Amphibians. U. C. Press, Berkeley, CA.

Welsh, H. H., Jr., and R. B. Bury. 2005. Del Norte salamander, Plethodon elongatus. in M. Lanno, editor. Status and Conservation of U. S. Amphibians. U. C. Press, Berkeley, CA.

Welsh, H. H., Jr., and N. E. Karraker. 2005. Southern torrent salamander, Rhyacotriton variegatus. in M. Lanno, editor. Status and Conservation of U. S. Amphibians. U. C. Press, Berkeley, CA.

Davic, R. D., and H. H. Welsh, Jr. 2004. On the ecological roles of salamanders. The Annual Review of Ecology, Evolution, and Systematics 35:405-434.

Welsh, H. H., Jr. 2004. Bookreview: LifePlace: Bioregional Thought and Practice. by Robert L. Thayer, Jr. The Quarterly Review of Biology 79:227-228.

Ollivier, L. M., and H. H. Welsh, Jr. 2003. Determining sex and life stage of Del Norte salamanders from external cues. Northwestern Naturalist 84:129-134.

Wheeler, C. A., H. H. Welsh, Jr., and L. L. Heise. 2003. Rana boylii oviposition Behavior. Herpetological Review 34:234.

Bettaso, J. B., H. H. Welsh, Jr., and B. D. Palmer. 2002. Northern red-legged frogs and endocrine-disrupting compounds (EDC's). FrogLog 52:1-2.

Welsh, H. H., Jr., and A. J. Lind. 2002. Multiscale habitat relationships of stream amphibians in the Klamath-Siskiyou Region of California and Oregon. Journal of Wildlife Management 66:581-602.

Welsh, Hartwell H., Jr.; Droege, Sam. 2001. A case for using plethodontid salamanders for monitoring biodiversity and ecosystem integrity of North American forests. Conservation Biology 15(3):558-569.

Welsh, Hartwell H., Jr.; Hodgson, Garth R.; Harvey, Bret C.; Roche, Maureen F. 2001. Distribution of juvenile coho salmon in relation to water temperatures in tributaries of the Mattole River, California. North American Journal of Fisheries Management 21(3):464-470.

Waters, Jeffrey R.; Zabel, Cynthia J.; McKelvey, Kevin S.; Welsh, Hartwell H., Jr. 2001. Vegetation patterns and abundances of amphibians and small mammals along small streams in a northwestern California watershed. Northwest Science 75(1):37-52.

Welsh, Hartwell H., Jr.; Roelofs, T.D.; Frissell, C.A. 2000. Aquatic ecosystems of the redwood region. Pages 165-199 in R.F. Noss, ed., The Redwood Forest: History, Ecology, and Conservation of the Coast Redwoods. Island Press, Covelo, California.

Welsh, Hartwell H., Jr.; Lind, A.J. 2000. Evidence of lingual-luring by an aquatic snake. Journal of Herpetology 34(1):67-74.

Thornburgh, D.A.; Noss, R.F.; Angelides, D.P.; Olson, C.M.; Euphrat, F.; Welsh, H.H., Jr. 2000. Managing redwoods. Pages 229-261 in R.F. Noss, ed., The Redwood Forest: History, Ecology, and Conservation of the Coast Redwoods. Island Press, Covelo, California.