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Integration of genetics and silviculture
Further development of options for increasing the quantity and quality of
wood production while considering other values requires a greater understanding
of the integration of genetics and silviculture.
Projects:
Genotype x silvicultural treatment interactions
Incorporating genetic effects into growth models
3.1 Genotype x silvicultural treatment interactions
The research activities of the Forest Genetics Team in the area of genotype
x silviculture interactions will focus primarily on continuing research on
the interaction of families with different levels of shade (Table 1). We have
initiated nursery studies to look at the response of ponderosa pine and hemlock
seedlings to different levels of shade using shade cloth, and these studies
will be completed. In cooperation with the PNW West-Side Silviculture Team,
we have also initiated a long-term field study on the Capitol Forest near Olympia
to consider the response of planted Douglas-fir, western hemlock, and western
red cedar to a range of different levels of overstory densities. This study
has a genetic component that considers the differential response of Douglas-fir
families. In addition to the shade studies, we will be working with Dick Miller,
formerly of the Silviculture Team, to complete a well-designed study of the
response of Douglas-fir families to fertilization at age 14.
Team Principal
Investigators: Brad St.Clair and Randy Johnson
Collaborators: Frank Sorensen
(retired PNW Genetics Team); Duane Ecker (Ochoco NF); Charlie Cartwright
(BC Ministry of Forests); Jeff DeBell (Washington DNR);
Leslie Brodie, David Marshall, Bob Curtis and Dean DeBell (PNW West-Side
Silviculture Team).
Benefits: Knowledge of family x silviculture interactions,
including interactions with shade and with fertilizer, will allow managers
to determine the value
of testing families from tree improvement programs under different silvicultural
regimes for selection and deployment to different regimes.
3.2 Incorporating
genetic effects into growth models
Research that provides knowledge of how to incorporate genetic effects into
growth models involves exploring the response to silvicultural treatments at
the level of the population, in this case, the improved population of selected
families. Most genetic trials involve individual tree plots and are not designed
to explore per-unit-area yields of a group of selected families in a large
plot. Realized gain trials using large block-plots look at levels of genetic
gains actually achieved from tree improvement activities, as compared to expected
genetic gains based on individual tree performance in progeny tests of all
families, not just the selected families. Few improved versus unimproved block-plot
realized genetic gain trials have been established throughout the world. The
Forest Genetics Team has been working with the Northwest Tree Improvement Cooperative
to establish such trials to be used to adjust existing growth models to take
into account the effects of genetic improvement (Table 1) (St.Clair 1993).
We are also working with the Stand Management Cooperative to establish trials
that consider the interactions of improved and unimproved populations with
different silvicultural treatments. And we have established a study to look
at the effects of intergenotypic competition by looking at the stand response
to deploying families in pure blocks versus in mixtures. These are long-term
studies; in the meantime, we will be working with the growth and yield modelers
to explore the effects on the outcomes from existing growth models using different
assumptions of the growth trajectories given what we know from progeny tests.
Team
Principal Investigators: Brad St.Clair and Randy Johnson
Collaborators: Keith
Jayawickrama (OSU), Greg Johnson (Willamette Industries) and other members
of the Northwest Tree Improvement Cooperative; David Marshall
(PNW Olympia Silviculture Team); David Briggs (University of Washington and
the Stand Management Cooperative)
Benefits: Knowledge of how genetically improved
populations grow given a different silvicultural options allow managers
to evaluate the value of and tradeoffs
among different management options.
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