02.PNW.B.1 -- Response of Native and Invasive Exotic Plants to Fire, Fire Recovery, and Fuel Reduction Treatments
in the Interior Pacific Northwest

 

CONTACT INFORMATION:
Team Leader: Catherine Parks (cparks01@fs.fed.us), Nan Vance (nvance@fs.fed.us)
Phone: 541-962-6531 (Parks), 541 750-7302 (Vance)

Research or Development Rationale:

In recent decades, non-native invasive plant species have increasingly supplanted native species on both disturbed and undisturbed sites in the interior Pacific Northwest. As non-native plants become more dominant in forest, grassland, and riparian plant communities, competitive simplification of native plant communities may result in loss of biodiversity, alteration of ecosystem function, and shifts in trajectories of plant succession. Although it is widely accepted that timber harvest and fire, both wild and prescribed, have direct and indirect influence on the establishment and persistence on non-native invasive plants, limited research has been conducted to test or support this concept. The knowledge base for management of non-native invasive plant species is largely derived from agricultural research, where techniques and treatments have been applied under controlled field conditions. Agricultural approaches can seldom be utilized in forests and rangelands. In addition, knowledge about the basic biology and ecology of many invasive plants is lacking, thus hampering effective and long-term control strategies.

The interior PNW is characterized by landscapes of mixed forests intergrading with temperate dry shrub steppe that have evolved under frequent fire intervals. These fires of varying intensity have resulted in a mosaic of plant communities, composed of species with different ecological amplitudes and adaptations to fire. These native communities, which generally undergo rapid post-fire recovery provide vegetative cover, and contribute to stability of montane slopes, and reduction of soil erosion potential. As a consequence of the transitional nature of this region, large, stand replacing fires that destroy canopy cover may convert forest types to more open grass types and barrens that favor aggressive exotic species usually associated with the dry steppe. It is not known if the displacement of native species by exotics increases the potential for erosion because exotics are less suitable for stabilization of soils. The specific effects of fire and fuel treatments, such as thinning, that cause changes in species dominance in these shifting mosaics are not well understood.

To add to current knowledge, we propose research with the following broad goals: 1) increasing knowledge on the basic ecology of selected invasive species and native forbs and grasses and their interaction with fire and disturbance, and 2) developing post-fire rehabilitation and restoration methods.

Research and Development Approach:

User needs/science gaps addressed: We will directly address the following science gaps identified under NFP Research Priority Biii: guidelines and practices for reestablishing native species and excluding invasive exotic plants on burned areas in long-term recovery of burned areas; conditions and requirements for developing seed sources of native plants; and methods for assessing, monitoring, and modeling post-fire vegetation recovery and plant invasions. Results will also contribute to science gaps identified under Research Priority Bii, particularly knowledge about the effects of wildfire and prescribed fire on habitat.

Managers addressing post-fire rehabilitation are giving high priority to achieving sufficient cover of native forbs, grasses and shrubs for long-term slope stability in addition to short term BAER treatments. Where evaluation indicates that native plant communities have not recovered, the reasons are ecologically complex. One hypothesis is that favorable conditions for the colonization and spread of invasive exotics confound the natural recovery of native plant communities. Increased cover and diversity of healthy plant communities may confer resistance to exotic invasions, but studies have been inconclusive. The interaction of invasive species with the their native plant neighbors needs to be studied if weed resistant plant communities are to be developed or maintained. Effective restoration requires identifying species that have characteristics of resistance to invasive exotic plants as well as soil stabilization properties. We plan to determine if increasing the cover of these species accomplishes both objectives.

Few studies have been conducted on responses of invasive plant species to wildfire and fuel reduction techniques. In forested ecosystems, considerable research has been conducted on fuels management and applied silvicultural problems; however, relatively few studies have addressed the effects of forest management on understory composition and diversity. In open grasslands and riparian areas the interest in increasing the diversity of native species and in building weed resistant plant communities is a high priority with restoration planners and commercial plant and seed suppliers. Basic information on plant species and community responses to wildfire and fuel reduction treatments (thinning, prescribed burning) are essential to resource managers in the assessment of ecological condition and trends, or for development of sound post-fire management strategies that enhance wildlife habitat and other components of biological diversity.
The proposed research seeks to address these science gaps through:

  1. Examining the biology and ecology of invasive species, particularly sulfur cinquefoil (Potentilla recta), cheat grass (Bromus tectorum) and yellow star thistle (Centaurea solstiliatis), in response to wildfire, prescribed burning, and rehabilitation treatments.
  2. Developing methods for reestablishing native species and rehabilitation of wildlife habitat (large game winter range) that are currently infested with exotic species. Adaptive management approaches and suggested treatment combinations for long-term control, monitoring, and assessment of invasive species will also be developed.
  3. Evaluating short and long-term effects of thinning, prescribed burning, and thinning + burning on the occurrence and abundance of invasive plant species, composition of forest understory plant communities, and seed production of dominant native shrub, grass and forb species.
  4. Analyzing existing permanent plot data for responses of vegetation to fire, particularly increased occurrence of invasive plant species and shifts in species dominance following wildfire in forest, grassland, and riparian communities.
  5. Analyzing the variation in plant communities in fire-created ponderosa pine/grassland mosaics with emphasis on fire's influence in shifting community types.
  6. Incorporating information from (5) evaluating a suite of dominant forb and grass species common to the dry Douglas-fir/ponderosa pine forest using the following criteria:
    a. effectiveness in immediate and long-term post-fire soil stabilization and rehabilitation,
    b. long-term use in restoring native plant communities,
    c. resistance to invasive exotics,
    d. providing local and regional commercial opportunities for native plant/seed suppliers.


Scientific Approach:

This research combines observational, experimental, and retrospective approaches.

Observational: We propose to characterize basic plant population biology and demography for the invasive perennial plant species, sulfur cinquefoil. The increase of sulfur cinquefoil in interior PNW has alerted managers to the urgent need for management recommendations, as well as basic plant biology information necessary to develop effective management strategies. Measurements will include seed production, dispersal, and seedling recruitment, and will be related to age of stand, and site variables (elevation, slope, aspect, soil type, plant community, land use, disturbance history). We will determine population age structure through examination of annual rings in the taproots using methods that have recently been developed in Europe (referred to 'herb-chronology'), and correlate age of populations and expansion patterns to wildfire and other disturbance events. We will also study the reproductive ecology to define its reproductive niche. The insect pollinator assemblage will be referenced, targeting infested areas where sulfur cinquefoil co-occurs with populations of native cinquefoil species.

Using GIS maps, aerial photographs, and plot data to examine dry forest/open land mosaic in the region, forest and open plant community types perpetuated by wildfire and topography will be identified and used to stratify the most commonly occurring native plant communities. Plant communities will be sampled in a replicated design for data that includes fire history and effect as well as other environmental attributes used to characterize the ecological land unit. This basic analysis will be used in the experimental studies to test suites of native plant species most suitable for post-fire restoration, weed resistance, and slope stabilization in forest, or open systems created by fire frequency and size.

Experimental: Effects of prescribed burning on the growth, density, seed bank dynamics, and seedling recruitment of invasive species, particularly sulfur cinquefoil and yellow star thistle, will be examined in experimental plots in at least two different study locations in northeast Oregon and one in west central Idaho. Responses of associated native species will also be monitored in treated plots. Rehabilitation treatments will include herbicide application and seeding with native grasses and forbs.

To evaluate native and exotic species for slope stability potential in the field we will assess ground cover development following wild fire in erosion susceptible sites. On runoff plots, life forms and life cycles of dominant plant species and soil surface conditions will be characterized. Experiments will include seeding plots (bare mineral soil) at different densities using non-native grasses, native plant species, and selected exotic invasive species. Summer precipitation, runoff and sedimentation will be measured. Temporal and spatial changes in plant species dominance will also be evaluated.

To evaluate the effects of fuel reduction treatments on invasive plant species, forest understory, and seed production of native species, we propose to augment ongoing research at three Fire and Fire Surrogate Study sites (Hungry Bob, near Enterprise, Oregon; Mission Creek, near Wenatchee, Washington, and Lubrecht Forest near Missoula, Montana). We will sample forest understory within the sites more intensively, and analyze baseline and vegetation changes through time (occurrence, abundance, native/exotic ratios) to test differences in occurrence and abundance of invasive species among treatments and between sites.

We also propose to measure seed production of dominant, native understory plant species of major habitat types on these three Fire and Fire Surrogate study sites. Seed production on thinned, burned and thinned and burned, and a control site will be collected for 3-5 years following fire or fire surrogate treatment. Native shrub, grass, and forb species will be evaluated for germination success and yield in agricultural multiplier environments. Seeds will then be re-introduced into different vegetation types. With this approach we expect to identify native indicator species that would be useful for short and long-term post fire rehabilitation efforts.

Retrospective: We will utilize the existing database from the USFS Ecology Program to select permanent plot locations that have been recently burned (within the last five years). Plot-level data will be analyzed for responses of vegetation to fire, particularly increased occurrence and expansion of invasive plant species following wildfire in forest, grassland, and riparian communities.

Development and technology transfer approach: We propose a workshop on the state of native plant seed source and availability and seed handling technology, and a regional infrastructure for R & D of native seed production and use. This provides a framework for studies that investigate which species are best suited for rehabilitation, i.e. which have the greatest potential to be used for seeding open, freshly disturbed areas, and which seed mixes are most compatible with early seral and established native plant communities over different soil types or conditions. This research, throughout its entirety, will be conducted with a broad range of university researchers, land owners, and land managers assuring the results are immediately available.

Description of products and how they will be used:

Products of this research will contribute fundamental knowledge about the biology and ecology of certain invasive species and native plants, their role and behavior in post-fire succession, their control and treatment in rehabilitation efforts, and interactions among species responses, fire, fuel reduction treatments. Interim results will be presented at regional and national scientific meetings, and local and regional workshops, and published through proceedings, newsletters, and PNW Station Research Notes. We anticipate the preparation and publication of a USFS General Technical Report (GTR) and journal article describing the techniques and preliminary findings of our 'herb chronology' research within the first two years of the project. Simple keys to distinguish invasive plant species from similar native congeners at different life stages will be published as user tools to assist in early, definitive detection of invasive species in assessment efforts. Final products targeted to the scientific community will be published as refereed journal articles, PNW Station Research Reports and/or GTRs; final products targeted to managerial user groups will be published as one or more GTRs, with guidelines and suggested protocols for rehabilitation efforts that control for the expansion of invasive plant species. Because most of the invasive species of concern are widespread throughout the U.S., we anticipate that the scientific results and guidelines will be used by ecologists and resource managers across multiple regions. An additional contribution of the project will be the establishment of study areas that can be repeatedly monitored over time for longer-term successional changes.

Research partners, roles and commitments: Oregon State University (through Steve Radosevich, Dept. of Forest Science) will partially support 1 post doctorate research associate and 1 graduate student. Oregon Department of Fish and Wildlife will provide available equipment/personnel to apply treatments on the Wildlife Management Areas.

Ed DePuit, PNW Station, Wenatchee Forestry Sciences Lab - collaboration on rehabilitation treatments.
Mike McInnis, Eastern Oregon University; plant/range/weed ecology; collaboration on weed biology research, interactions of invasive and native plant species;
Linda Wilson, University of Idaho; modeling population biology, demography, and projected expansion.
Charles Johnson, NFS Area Ecologist,- plant ecology, synthesis of existing databases on natives.
Marty Vavra, Eastern Oregon Agricultural Research Center, ungulate/range ecology.
Rocky Mountain Elk Foundation; ungulate ecology; rehabilitation of winter range
John Kie, LaGrande Lab, PNW Research Station; wildlife ecology
Dr. Peter Bernhardt, Research Fellow, Missouri Botanical Gardens, St. Louis (joint venture) pollination ecology of sulfur cinquefoil
NRCS Plant Materials Center, Corvallis, Oregon, collaboration in seed and cultivation of plant material
JH Stone Nursery, Central Point, Oregon, collaboration in producing plant material.
Fritz Sweingruber, Swiss Federal Research Institute for Forests, Birmensdorf, Switzerland

Potential Users:
Center for Invasive Plant Management, Montana State University, Bozeman, Montana
National Forest Systems in R6, R4, and R1, USDI Bureau of Land Management
The Nature Conservancy, Private Land Owners, Commercial native plant and seed suppliers in the region.
Forest and Weed Scientists, State and Private Foresters
Oregon, Washington, and Idaho Departments of Agriculture, Noxious Weed Programs
Tri-County Weed Management Area, northeast Oregon
Regional and forest botanists and plant ecologists in R1,R4, and R6

Linkages and integration: The proposed research is described as a stand-alone project. However, if funded, we intend to integrate research with existing (and proposed) projects across the West.

Co-investigators: Dana Perkins, Research Ecologist, PNW Station, La Grande, and Kate Dwire, Research Riparian Ecologist, RM Station, Laramie.

Lead Research Work Unit: Managing Natural Disturbance Team, MDR Program, La Grande, and Biology and Culture of Forest Plants Team, RMP Program, Corvallis, Oregon.

Lead Research Station: PNW Research Station