SPECIES: Populus deltoides



Photos by:     

Bill Cook, MI State Univ., www.forestryimages.org. Eastern cottonwood. W.L. Wagner @ USDA-NRCS PLANTS Database. Rio Grande cottonwood.

Taylor, Jennifer L. 2001. Populus deltoides. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/ [].


Populus deltoides var. occidentalis Rydb.
   = P. d. ssp. monilifera (Ait.) Eckenwalder [126,128]
P. fremontii var. wislizeni S. Wats
   = P. d. ssp. wislizeni (S. Wats.) Dorn [56,126,128,170]
P. monilifera Ait.
   = P. d. ssp. monilifera (Ait.) Eckenwalder [128]
P. sargentii Dode
   = P. d. ssp. monilifera (Ait.) Eckenwalder [88,126,128]
P. wislizeni (S. Wats.) Sarg.
   = P. d. ssp. wislizeni (S. Wats.) Dorn [56,126,128,170]


eastern cottonwood
plains cottonwood
Rio Grande cottonwood
plains poplar

The currently accepted name of eastern cottonwood is Populus deltoides Bartr. ex Marsh. (Salicaceae) [56,128]. It is in the Aigeiros section of Populus [86]. Recognized infrataxa are as follows:

Populus deltoides ssp. deltoides Bartr. ex Marsh.  eastern cottonwood 
Populus deltoides ssp. monilifera (Ait.) Eckenwalder    plains cottonwood [48,126,128]
Populus deltoides ssp. wislizeni (S. Wats.) Eckenwalder   Rio Grande cottonwood [48,56,126,128]

In this species summary eastern cottonwood refers to the species (Populus deltoides) unless otherwise noted as P. d. ssp. deltoides. Plains cottonwood and Rio Grande cottonwood refer to P. d. ssp. monilifera and P. d. ssp. wislizeni, respectively.

Eastern cottonwood produces several hybrids:

Carolina poplar (P. × canadensis Moench); from hybridization with Lombardy cottonwood (P. nigra) [56,57,88]
lanceleaf cottonwood (P. × acuminata Rydb.); afrom hybridization with narrowleaf cottonwood (P. angustifolia) [47,88]
balm-of-Gilead (P. × jackii Sarg.); from hybridization with balsam poplar (P. balsamifera) [47,57,65,88]


No special status

In Texas, the cottonwood-tallgrass series is listed as "imperiled globally, very rare, 6 to 20 occurrences (endangered throughout range); imperiled in Texas, very rare, vulnerable to extirpation, 6 to 20 occurrences" [206].


SPECIES: Populus deltoides

Eastern cottonwood occurs from Alberta east to Quebec and south to Florida, Texas, Arizona, and northern Mexico [128]. Populus deltoides ssp. deltoides occurs from the Gulf of Mexico north along the Atlantic coast to Maine and Quebec; along the Mississippi River to Illinois and Ohio; and westward to Texas and Oklahoma [55,85]. Plains cottonwood ranges from Alberta east to Quebec and south to Rocky Mountain foothills, the Great Plains, Pennsylvania, the Texas panhandle, and New Mexico [55,88,147,173,194]. Rio Grande cottonwood occurs from northeastern Utah and northwestern Colorado through the upper Rio Grande and Colorado Plateau regions to southern Arizona and New Mexico, southwestern Texas, and northern Mexico [55,56,147]. The PLANTS database provides distributional maps of eastern cottonwood, Populus deltoides ssp. deltoides, plains cottonwood, and Rio Grande cottonwood.

FRES12 Longleaf-slash pine
FRES13 Loblolly-shortleaf pine
FRES14 Oak-pine
FRES15 Oak-hickory
FRES16 Oak-gum-cypress
FRES17 Elm-ash-cottonwood
FRES18 Maple-beech-birch
FRES19 Aspen-birch
FRES21 Ponderosa pine
FRES29 Sagebrush
FRES30 Desert shrub
FRES32 Texas savanna
FRES33 Southwestern shrubsteppe
FRES35 Pinyon-juniper
FRES36 Mountain grasslands
FRES38 Plains grasslands
FRES39 Prairie
FRES40 Desert grasslands
FRES41 Wet grasslands


1 Northern Pacific Border
2 Cascade Mountains
3 Southern Pacific Border
4 Sierra Mountains
5 Columbia Plateau
6 Upper Basin and Range
7 Lower Basin and Range
8 Northern Rocky Mountains
9 Middle Rocky Mountains
10 Wyoming Basin
11 Southern Rocky Mountains
12 Colorado Plateau
13 Rocky Mountain Piedmont
14 Great Plains
15 Black Hills Uplift
16 Upper Missouri Basin and Broken Lands

K011 Western ponderosa forest
K016 Eastern ponderosa forest
K017 Black Hills pine forest
K023 Juniper-pinyon woodland
K027 Mesquite bosques
K038 Great Basin sagebrush
K055 Sagebrush steppe
K056 Wheatgrass-needlegrass shrubsteppe
K063 Foothills prairie
K064 Grama-needlegrass-wheatgrass
K065 Grama-buffalo grass
K066 Wheatgrass-needlegrass
K067 Wheatgrass-bluestem-needlegrass
K068 Wheatgrass-grama-buffalo grass
K069 Bluestem-grama prairie
K070 Sandsage-bluestem prairie
K074 Bluestem prairie
K081 Oak savanna
K085 Mesquite-buffalo grass
K088 Fayette prairie
K089 Black Belt
K098 Northern floodplain forest
K100 Oak-hickory forest
K101 Elm-ash forest
K104 Appalachian oak forest
K106 Northern hardwoods
K109 Transition between K104 and K106
K111 Oak-hickory-pine
K112 Southern mixed forest
K113 Southern floodplain forest

39 Black ash-American elm-red maple
42 Bur oak
46 Eastern redcedar
61 River birch-sycamore
62 Silver maple-American elm
63 Cottonwood
92 Sweetgum-willow oak
93 Sugarberry-American elm-green ash
94 Sycamore-sweetgum-American elm
95 Black willow
109 Hawthorn
220 Rocky Mountain juniper
235 Cottonwood-willow
236 Bur oak
242 Mesquite

203 Riparian woodland
314 Big sagebrush-bluebunch wheatgrass
403 Wyoming big sagebrush
408 Other sagebrush types
412 Juniper-pinyon woodland
421 Chokecherry-serviceberry-rose
422 Riparian
501 Saltbush-greasewood
504 Juniper-pinyon pine woodland
601 Bluestem prairie
602 Bluestem-prairie sandreed
604 Bluestem-grama prairie
609 Wheatgrass-grama
610 Wheatgrass
611 Blue grama-buffalo grass
612 Sagebrush-grass
704 Blue grama-western wheatgrass
708 Bluestem-dropseed
709 Bluestem-grama
710 Bluestem prairie
715 Grama-buffalo grass
727 Mesquite-buffalo grass
729 Mesquite
801 Savanna
802 Missouri prairie
805 Riparian

Eastern cottonwood often occurs as a dominant or codominant component of floodplain and bottomland hardwood forests [28,42,70,112,158]. It is a principal species in riverfront forests in the eastern United States [150]. The maintenance of eastern cottonwood-dominated stands depends on periodic flooding [96,155,229]. Most of these riparian areas tend to be in early successional stages and are composed chiefly of scrub willows (Salix spp.) interspersed with occasional eastern cottonwood stands [155].

Trees: Throughout its range, eastern cottonwood can grow in pure stands [145], but more often grows in mixed stands [10,28]. Common tree associates include hackberry (Celtis occidentalis) [26,30,90,111,141], sugarberry (C. laevigata) [111,141,150,161], boxelder (Acer negundo) [10,30,111,114,131,141,161,166], silver maple (A. saccharinum) [2,8,10,51,102,111,141,202,221], sycamore (Platanus occidentalis) [2,26,114,131,141,150,161,196], American elm (Ulmus americana) [8,26,62,99,111,114,141,150,161,166,202], green ash (Fraxinus pennsylvanica) [2,3,8,62,70,99,111,141,150], sandbar willow (Salix exigua) [8,166,234], and bur oak (Quercus macrocarpa) [118,141].

Other tree species found with eastern cottonwood include swamp white oak (Quercus bicolor), pin oak (Q. palustris) [90,141], white mulberry (Morus alba), black cherry (Prunus serotina),  chestnut oak (Quercus prinus), black oak (Q. velutina), sassafras (Sassafras albidum), tree-of-heaven (Ailanthus altissima), bigtooth aspen (Populus grandidentata), red elderberry (Sambucus racemosa ssp. pubens) [90], American hornbeam (Carpinus caroliniana), bitternut hickory (C. cordiformis) [114,141], white ash (Fraxinus americana) [114,131], basswood (Tilia americana) [99,114], balsam poplar (Populus balsamifera), pussy willow (Salix discolor), speckled alder (Alnus rugosa) [8], black willow (Salix nigra) [10,62,161,166], Shumard oak (Quercus shumardii) [141,219], post oak (Q. stellata), cherrybark oak (Q. pagoda), northern red oak (Q. rubra), Kentucky coffeetree (Gymnocladus dioicus), shagbark hickory (Carya ovata),  mockernut hickory (C. tomentosa), shellbark hickory (C. lacinios), pignut hickory (C. glabra), river birch (Betula nigra), red maple (Acer rubrum), sweetgum (Liquidambar styraciflua), eastern redbud (Cercis canadensis), pawpaw (Asimina triloba), honey-locust (Gleditsia triacanthos) [141], hawthorns (Crataegus spp.), common persimmon (Diospyros virginiana) [141,161], black walnut (Juglans nigra) [3,123,141], chinkapin oak (Q. muehlenbergii), white oak (Q. alba), [118], peachleaf willow (Salix amygdaloides) [99,221], pecan (Carya illinoensis) [26,150,161,166], French tamarisk (Tamarix gallica) [166], eastern swampprivet (Forestiera acuminata), red mulberry (Morus rubra), Chinaberrytree (Melia azedarach), water hickory (Carya aquatica), buttonbush (Cephalanthus occidentalis) [161], gum bully (Sideroxylon lanuginosum) [30], and cedar elm (Ulmus crassifolia) [30,161].

Shrubs: Some common shrubs associated with eastern cottonwood are winged burning bush (Euonymus alata), northern spicebush (Lindera benzoin), black tupelo (Nyssa sylvatica), multiflora rose (Rosa multiflora), southern arrowwood (Viburnum dentatum), blackhaw (V. prunifolium) [90], sand shinnery oak (Quercus havardii), sand sagebrush (Artemisia filifolia) [206], and common ninebark (Physocarpus opulifolius) [8].

Graminoids: Graminoid species found in the understory of eastern cottonwood stands include streambank wheatgrass (Elymus lanceolatus ssp. psammophilus), sanddune sandbur (Cenchrus tribuloides) [234], Canada wildrye (Elymus canadensis) [166,234], switchgrass (Panicum virgatum) [206], Japanese brome (Bromus japonicus), Bermuda grass (Cynodon dactylon), johnsongrass (Sorghum halepense) [166], big bluestem (Andropogon gerardii var. gerardii), Indiangrass (Sorghastrum nutans), alkali sacaton (Sporobolus airoides), and eastern gamagrass (Tripsacum dactyloides) [206].

Forbs and vines: Several forbs and vines are found in eastern cottonwood stands. Some common species are common moonseed (Menispermum canadense), Asian bittersweet (Celastrus orbiculatus), privet (Ligustrum spp.) [90], poison-ivy (Toxicodendron radicans), [90,111,166,234], trumpet creeper (Campsis radicans), oneseed bur cucumber (Sicyos angulatus) [111], peppervine (Ampelopsis arborea), grape (Vitis spp.), Rubus species [111,166], Smilax species, Helianthus species, American pokeweed (Phytolacca americana) [26], Virginia creeper (Parthenocissus quinquefolia) [111,166], goldenrod (Solidago spp.), saw greenbrier (Smilax bona-nox), great ragweed (Ambrosia trifida), Canadian horseweed (Conyza canadensis) [166], white sweetclover (Melilotus alba), roughleaf dogwood (Cornus drummondii), coltsfoot (Tussilago farfara), and American searocket (Cakile edentula) [234].

Classifications systems listing eastern cottonwood is an indicator species or as a dominant component of community types, habitat types, plant associations, or riparian site types are as follows:

Illinois [2]
Massachusetts [131]
Missouri [51]
New York [174]
Tennessee [187,192,193]
Quebec [43]

Plains cottonwood:
Trees: Common tree associates of plains cottonwood include boxelder [4,9,12,25,39,61,74,92,105,124,130,143], green ash [4,9,12,19,25,39,54,61,74,83,84,92,104,124,130], American elm [9,12,19,25,39,61,104,105,124,130,143], chokecherry (Prunus virginiana) [19,54,92,130,143,163,208], peachleaf willow [12,20,25,36,61,95,124,130,143], red-osier dogwood (Cornus sericea) [92,95,130,143,208], Russian-olive (Elaeagnus angustifolia) [36,95], willows (Salix spp.) [4,17,61,92,139,163,208], bur oak [19,25,61,104,105,124,130,143,173], silver maple [4,61], hackberry [12,25,61], black willow [42,143], and sandbar willow [4,12,25,143,175].

Other trees found with plains cottonwood include silver buffaloberry (Shepherdia argentea) [54,139], river willow (Salix fluviatilis) [20], black cottonwood (Populus balsamifera ssp. trichocarpa) [96], narrowleaf cottonwood [95,96], Rocky Mountain juniper (Juniperus scopulorum) [83,84,163], Cornus species [61], black willow [42,143], red elderberry [143], fivestamen tamarisk (Tamarix chinensis) [17], red mulberry [12,105], basswood [25,105], northern red oak, chinkapin oak, bitternut hickory, slippery elm (Ulmus rubra), black oak (Quercus velutina), white ash, eastern redbud, black cherry [105], American plum (Prunus americana), eastern redcedar (Juniperus virginiana) [19], eastern hophornbeam (Ostrya virginiana) [25,105], eastern redcedar, black walnut, Kentucky coffeetree, slippery elm, and rock elm (Ulmus thomasii) [25].

Shrubs: Shrubs associated with plains cottonwood are western snowberry (Symphoricarpos occidentalis) [4,20,54,83,84,92,130,143,163,175], Wood's rose (Rosa woodsii) [20,83,84,92,95], plains silver sagebrush (Artemisia cana ssp. cana), [20,50], golden currant (Ribes aureum) [92], Rocky Mountain maple (Acer glabrum)  [208], Saskatoon serviceberry (Amelanchier alnifolia)  [130,208], Douglas hawthorn (Crataegus douglasii) [208], American hazel (Corylus americana) [105,143], American black currant (Ribes americanum), alderleaf buckthorn (Rhamnus alnifolia), smooth rose (Rosa blanda) [143], common reed (Phragmites australis) [175], desert false indigo (Amorpha fruticosa) [4], toughleaf dogwood (Cornus asperifolia), red raspberry (Rubus idaeus), common cocklebur (Xanthium strumarium), eastern wahoo (Euonymus atrourpureua), smooth sumac (Rhus glabra) [105], and skunkbush sumac (R. trilobata) [107].

Graminoids: Graminoid species found in the understory of plains cottonwood stands include blue grama (Bouteloua gracilis) [4,50], cheatgrass (Bromus tectorum) [50], Canada wildrye [20,36,83,92,95,175], prairie cordgrass (Spartina pectinata) [175], sand dropseed (Sporobolus cryptandrus), buffalo grass (Buchloe dactyloides) [4], barnyard grass (Echinochloa crus-galli) [20,143], western wheatgrass (Pascopyrum smithii) [20,36], prairie sandreed (Calamovilfa longifolia) [20,107], smooth brome (Bromus inermis) [20,36,92,95], quackgrass (Elytrigia repens), green muhly (Muhlenbergia racemosa) [20,92,95], hairy wildrye (Elymus villosusi), creeping bentgrass (Agrostis stolonifera), timothy (Phleum pratense) [92,95], Kentucky bluegrass (Poa pratensis) [36,92,95,107,143], sedges (Carex spp.), Bromus species, Scirpus species [39], Virginia wildrye (Elymus virginicus) [36], Assiniboia sedge (Carex assiniboinensis), and slender wheatgrass (Elymus trachycaulus) [143].

Forbs and vines: Forbs and vines found in plains cottonwood stands are Virginia creeper [20,92,95,105,124], western poison-ivy (Toxicodendron rydbergii) [20,92,95], peppervine [92], longroot smartweed (Polygonum amphibium), golden dock (Rumex maritimus), riverbank grape (Vitis riparia), Louisiana sagewort (Artemisia ludoviciana) [20], white sweetclover [20,83,84,95,107,175], wild licorice (Glycrrhiza lepidota) [20,95], yellow sweetclover (Melilotus officinalis) [95], American bittersweet (Celastrus scandens), frost grape (Vitis vulpina) [105,124,175], western white clematis (Clematis ligusticifolia) [124], poison-ivy [36,143,175], woodbine, American vetch (Vicia americana), Canada thistle (Cirsium arvense), common milkweed (Asclepias syriaca) [36], starry Solomon's seal (Maianthemum stellatum) [83,84,143], veiny meadowrue (Thalictrum venulosum), purple meadowrue (T. dasycarpum) [83,84], smooth Solomon's seal (Polygonatum biflorum var. commutatum), California nettle (Urtica dioica), heartleaf four o' clock (Mirabilis nyctaginea), wild sarsaparilla (Aralia nudicaulis)], creeping violet (Viola canadensis var. rugulosa), fireweed (Epilobium angustifolium), moist sowthistle (Sonchus arvensis ssp. ulginosus), smooth aster (Aster laevis), white panicle aster (Symphotrichum lanceolatum ssp. lanceolatum), wild mint (Mentha canadensis), common hop (Humulus americanus), Blue Ridge carrionflower (Smilax lasioneura), hedge false bindweed (Calystegia sepium) [143], lambsquarters (Chenopodium album), poison hemlock (Conium maculatum), curly dock (Rumex crispus) [175], common sunflower (Helianthus annuus), Chenopodium species, and prickly Russian-thistle (Salsola tragus) [4].

Classifications systems listing plains cottonwood is an indicator species or as a dominant component of community types, habitat types, plant associations, riparian site types, or dominance types are as follows:

Montana [92,93,94,95]
North Dakota [83,84,223]
Wyoming [162,207]
Manitoba [143]

Rio grade cottonwood:
Common tree associates of Rio Grande cottonwood include peachleaf willow [73,113], hybrid crack willow (Salix x rubens) [73], Russian-olive, Goodding willow (Salix gooddingii) [59,113], saltcedar (Tamarix ramosissima) [59], and sandbar willow (Salix exigua) [113]. Understory species include mule's fat (Baccharis glutinosa), desert false indigo, stretchberry (Forestiera neomexicana), screwbean mesquite (Prosopis pubescens), and Torrey wolfberry (Lycium torreyi) [59,113].

Classifications systems listing Rio Grande cottonwood is an indicator species or as a dominant component of community types, habitat types, plant associations, or riparian site types are as follows:

Arizona [204]
New Mexico [53,116,204]
Texas [46,206]


SPECIES: Populus deltoides

The wood of eastern cottonwood is moderately light in weight, rather soft, and relatively weak in bending and compression [47,132,145,210]. It is uniform in texture and usually straight grained [28,47,210]. Primary wood products include lumber [28,132,210,212,217], veneer [28,47,132,145,151,210,217], plywood [132], excelsior [28,47,151,210], fiberboard [28,45,151,212], paper pulp [28,45,132,151,212,217], sawtimber [47], and pulpwood [47,132,210]. Finished wood products include pallets, crates [28,145,151], furniture [145], and food containers [132]. Eastern cottonwood is slightly to nonresistant to heartwood decay [210].

Eastern cottonwood is a valuable timber species [28,159]. It is used as a short-rotation intensive culture species in the southern United States [47,160] and Canada [144], and is highly suitable for plantation management [75,134,145].

Plains cottonwood has similar wood characteristics [132,173], but is not considered to be commercially valuable [132]. The wood is not durable when exposed to soil and other moist conditions. It is used for rough construction lumber, temporary fence posts, corral poles, fuel, veneer, boxes, plywood, excelsior, and wood pulp [173].

In the northern Great Plains, eastern cottonwoods are a component of riparian forests and moist woodlands that provide critical habitat for many wildlife species [9,15,107,119,187,200]. These woodland areas may constitute up to 50% of the habitat for deer and 70% of the habitat for sharp-tailed grouse throughout much of the Great Plains. Domestic livestock use these communities for shade, forage, and water in the summer, and for thermal cover in the winter [15]. Eastern cottonwood has been classified as having fair value for all wildlife, songbirds, upland game birds, fur and game mammals [32].

The bark and leaves of eastern cottonwood seedlings and saplings are eaten by field mice, rabbits, deer, and domestic livestock [9,28,122,145]. Wesley and others [226] observed the use of eastern cottonwood plantations in Arkansas and Mississippi by wild turkeys for courtship, prenesting, nesting, and poultry rearing. Eastern cottonwood plantations were used by white-tailed does, rabbits, and northern bobwhite more than surrounding natural stands.

Plains cottonwood stands provide habitat for 82% of all bird species breeding in northeastern Colorado [181]. These forests provide roosting and nesting sites [21,80,91,107,153,167], feeding sites [63,107,116,205], and nest material for several bird species [80]. Beavers use the wood of plains cottonwood for food and for buildings dams and lodges [92]. The plains cottonwood/red-osier dogwood community provides thermal cover, debris recruitment, and streamside stability for fishes [92,173]. Plains cottonwood is eaten by prairie porcupines [98] and is the most important browse species for mule deer in the fall [148].

Evans and Dietz [63] found plains cottonwood was the least palatable to sage grouse of all trial foods. The palatability of plains cottonwood has been rated as follows [49]:

Cattle Poor Poor Fair Fair
Domestic sheep Poor Fair Fair Fair
Horse Poor Poor Fair Fair

In Colorado, the palatability of Rio Grande cottonwood has been rated poor for cattle, domestic sheep, and horses [49]:

Plains and Rio Grande cottonwood have been rated as fair in energy and protein value. The gross energy value of plains cottonwood is 5.385%, crude protein 5.4% (oven-dried weight), and metabolizable energy 2.686% (air dried) [63]. The wildlife food value of plains cottonwood in 3 western states has been rated as follows [49]:

Elk Poor ---- Good
Mule deer Poor Fair Good
White-tailed deer Fair Fair Good
Pronghorn ---- Fair Poor
Upland game birds ---- Fair Poor
Waterfowl ---- ---- Poor
Small non-game birds ---- Poor Fair
Small mammals ---- ---- Good

Eastern cottonwoods provide a nesting place for white-throated sparrows and myrtle warblers [187]. They provide roost sites for Rio Grande turkeys [14] and nursery colonies for the Indiana bat [24]. In North Dakota, eastern cottonwoods provide important night roosting cover in winter for the greater prairie-chicken and sharp-tailed grouse [146]. Eastern screech-owls are found in plains cottonwood dominated riparian woodlands east of the Continental Divide [69]. Plains cottonwoods are a very important nesting substrate for nesting raptors. Plains cottonwoods are used for nesting by golden and bald eagles, several hawk species, Lewis' and red-headed woodpeckers, and other cavity nesters [17,167,179,180].

The only natural habitat for fox squirrels is plains cottonwood bottomlands.  Plains cottonwood stands in Colorado are used by fox squirrels for nesting and feeding [232]. Plains cottonwood forests provide nesting sites for eagles, hawks, and other birds [17,69,167]. The cover value of plains cottonwood for some wildlife species has been rated as follows [49]:

Elk ---- Fair ---- Good
Mule deer ---- Fair Fair Good
White-tailed deer Fair Good Good Good
Pronghorn ---- ---- Poor Poor
Upland game birds Poor Fair Poor Good
Waterfowl ---- ---- ---- Poor
Small non-game birds Good Fair Good Good
Small mammals Good Poor ---- Good

The cover value of Rio Grande cottonwood in Colorado has been rated as follows [49]:

White-tailed deer Fair
Upland game birds Poor
Small non-game birds Good
Small mammals Good

Eastern cottonwood is well suited for revegetating disturbed riparian sites and has also been used extensively in the reclamation of strip-mined lands [27,157,217]. Eastern cottonwood (P. d. ssp. deltoides) has been planted successfully on mine spoils in Ohio both in pure stands and in mixture with black locust (Robinia pseudoacacia) [140]. The extensive root system holds streambanks in place [220], is effective in shoreline protection, and revegetating eroded stream channels [101]. Eastern cottonwood can be used as living dams for erosion and flood control work [145].

Eastern cottonwood may establish on suitable sites through natural seedfall [138,177,216], or it may be established by cuttings [47,68,133]. The average length of cuttings in the Pacific Northwest and the southern United States is about 20 inches (50 cm), while 8- to 12-inch (20-30 cm) cuttings are typical in the northern United States and Canada. However, cuttings of 8 feet (2.4 m) or more planted in 3-foot (1 m) deep holes have advantages over standard 20-inch (50 cm) cuttings. These advantages include less intensive site preparation requirements, a reduced need for browsing protection, and less intensive weed control [133]. In general, cuttings should be longer where upper soil moisture is limiting [47]. In Ohio, eastern cottonwood had better growth and survival when planted on loamy and clayey soils of previously coal strip-mined lands [140].

The growth of young cottonwood seedlings on favorable sites is rapid, but the plants must be kept free of competing vegetation to survive [34,132]. Browsing and trampling by wildlife and domestic animals must also be controlled for successful growth [47]. Whether revegetating by seeding or cuttings, native stock should be selected if available, since significant geographic variation exists in growth rate, drought resistance, wood characteristics, and sprouting ability [41,169].

Plains cottonwood was planted on surface-mined lands in Indiana 1928-1975 [27]. Rio Grande cottonwood is recommended for planting in the western Great Plains and desert southwest. Plains cottonwood is recommended for the northern Great Plains and western United States [33].

Eastern cottonwood is relatively drought resistant [47] and has been used extensively in shelterbelt and windbreak plantings in the Great Plains region of the United States and Canada [35,173,178,209]. Hybrid clones bred for improved winter hardiness and resistance to insects and diseases are commonly used in such plantings [47]. Eastern cottonwood was introduced to Victoria, Australia, and is recommended for use in fire shelterbelts [190].

Plains cottonwood, the "Pioneer Tree of the Plains," is often the only tree found in the western United States. It is a sure sign of water and welcome shade [31,47,173]. During severe winters saplings were used as horse and cattle feed by Native Americans and early settlers [9,47,81]. Native Americans used the roots to start fires [31] and used smaller trees for lodge poles and travois. The teepee pattern is supposedly patterned after the deltoid leaf shape [173]. The Teton Dakota ate the inner bark and the Omaha used it to make the Sacred Pole. Nebraska tribe children made toys with the leaves and made gum and play jewelry from the fruits [81].

Plains cottonwood grows into an effective windbreak in 15 to 20 years, reaching 40 to 50 feet (12.2-15.2 m). It is recommended for planting in shelterbelts only if irrigated, on wetter sites, or in rows near the center of the shelterbelt [78,79].

The wood of Rio Grande cottonwood was used by the Navajo for firewood, fence posts, cradles, tinderboxes, wooden tubes of bellows, dolls, and images for ceremonies. Chewing gum was made from the sap or the catkins mixed with animal fat [60].

Eastern cottonwood seedlings are highly susceptible to grazing and trampling damage from wildlife and domestic livestock. Control measures generally must be taken, particularly on plantations, for successful establishment and growth [9,40,47,145]. These can include the selection of certain clones that may be somewhat less palatable, weed control, use of repellent materials, fencing, and direct control of the animals [47]. Eastern cottonwood should not be planted next to European alder (Alnus glutinosa) [168] or Siberian elm (Ulmus pumila) in shelterbelts, because it will be shaded and mortality occurs within 10 years [191].

Strips or buffer zones of eastern cottonwood stands immediately adjacent to streams and rivers are effective for erosion control [96].

In the Great Plains region, recruitment and survival of plains cottonwood has been adversely affected by the construction of dams and reservoirs. Changes in magnitude and frequency of floods, rates of sedimentation, and rates of meander migration contribute to the reduction of suitable recruitment sites [23,124]. Periodic large releases of reservoir water to simulate natural flooding are recommended to ensure vigorous recruitment and growth of cottonwood forests on prairie river floodplains [23]. Flooding also decreases litter accumulation and can reduce the threat of fire [59]. In Colorado, moderately high flows that occur every 5 years are required to create the new point bars where plains cottonwood establishes [72]. Seedlings will establish close to the edges of river channels, but will probably not survive future ice jams and high discharges. The long-term survival of seedlings established during flood-free periods is greater the higher above stream channel they are established [179]. See black cottonwood for further information on the effects of watercourse damming and stream diversion on plains and other cottonwoods.

Disease and insect pests that affect eastern cottonwood have been described by several authors [28,44,156,173,195].

Russian-olive and saltcedar have invaded many riparian woodlands across the Great Plains and southwestern United States dominated by cottonwoods (Populus spp.) and willows. The invaders have displaced the native vegetation, taken up water, and increased fire frequency [137,185,197]. Russian-olive and saltcedar provide habitat for some wildlife species [137,185]. However, the loss of larger trees, especially eastern cottonwoods, has led to a decrease in habitat for cavity-nesting birds [185].


SPECIES: Populus deltoides

Plains cottonwood photo by:  

J.S. Peterson@USDA-NRCS PLANTS Database

Eastern cottonwood is a native, deciduous bottomland hardwood [68,121,132,212,225]. Height ranges from 36 to 190 feet (11-57.9 m) [37,47,56,67,120,132,150,151,224]. At maturity (approximately 35 years) [132], diameter at breast height ranges from 10.7 inches to more than 6 feet (27.2-182.9 cm) [7,47,67,132,150,224]. In open areas, eastern cottonwood typically has a large trunk that divides into branches near its base and ascends to form a wide, spreading crown [47,100]. In closed stands, it tends to have a tall, straight, and relatively branch-free bole with a small rounded crown [9]. Life expectancy is approximately 100 to 200 years [9,120,142]. It is dioecious. Female catkins range from 2 to 5.1 inches (5-13 cm) long, and fruit capsules are 0.3 to 0.6 inch (.8-1.5 cm) long [56]. The bark is thick and deeply furrowed with wide, flat ridges [56,199]. The rooting depth averages 100 inches (254 cm) [97], and mature stands can reach 117.6 to 196.8 inches (298.7-499.9 cm) rooting depth [22]. 

Eastern cottonwood is drought tolerant [47,77]. It has been classed as moderately tolerant to water-logged soils [106,110] and is tolerant of short-term inundation [145]. Eastern cottonwood tolerates periodic flooding from January through April, but mortality and growth due to flooding depend on how many events per year, season of year, age class, duration, and depth [89,106,149]. A study using cuttings found that eastern cottonwood survived less than 16 days of complete submergence [109]. Roots die when soaked for more than 1 month and adventitious roots form from dormant buds in the main trunk [110,220].

Plains cottonwood is a quick growing, short-lived deciduous tree [4]. It is smaller than P. d. ssp. deltoides [173] and grows from 10 to 98.4 feet (3-30 m) [4,36,58,124,130,139,173,195,207,223]. It has a diameter at breast height of 5 to 78.7 inches (12.7-200 cm) [4,58,107,124,130,139,173,207]. It is considered the fastest-growing tree in the Great Plains. The life expectancy is about 90 years [173].

Staminate catkins of plains cottonwood range in size from 2 to 3.5 inches (5-9 cm), pistillate catkins are 5.9 to 7.9 inches (15-20 cm) long, and fruits are 0.4 inch (1 cm) in length [173]. Plains cottonwood is susceptible to drought except where water tables are high [4] and drought induced mortality can be high [173]. The average rooting depth is 10 feet (3.1 m), and the longest roots can reach 75 feet (22.9 m) [231].

Rio Grande cottonwood ranges in height from 26.2 to 65.6 feet (8-20 m) [56,59]. It has catkins with pedicels 0.2 to 0.6 inch (.5-1.5 cm) long [56].


Eastern cottonwood regenerates sexually and vegetatively.

Sexual reproduction: Large seed crops (25 to 28 million seeds/tree/year) [9,22,129] dispersed by wind and water over long distances, are generally produced annually once the trees reach 10 to 15 years of age [9,22,47,132,142,158,173,221,225].

There is no seed dormancy in eastern cottonwood [227,228]. Germination occurs as soon as seeds arrive on a seedbed that is moist, free of vegetation, and in full sunlight [9,23,67,132,173,181]. Seeds are highly viable at dispersal [66,124], but viability decreases rapidly in the absence of a suitable germination environment [66]. Under natural conditions, seeds remain viable for 1 to 2 weeks [22]. Suitable recruitment sites occur naturally as a result of spring flooding [156,173] and scouring by ice [9]. Receding floodwaters leave freshly deposited, exposed alluvium, and seed germination along prairie river floodplains often occurs exclusively on these sites [124,229]. Exposed soil is essential, as young seedlings do not compete well with overtopping vegetation [34,124,171].

Vegetative reproduction: Cottonwood species (Populus spp.) reproduce vegetatively by sprouting from stumps and root crowns, and by the formation of suckers (adventitious shoots on roots ) [47,176,229]. There is asexual reproduction from broken limbs (flood training) and crown breakage [22]. The ability of cottonwoods to sprout declines with age [173].  Cottonwoods in the Aigeiros section of Populus, including eastern cottonwood, do not sprout as readily as cottonwoods in section Tacamahaca. Eastern cottonwood sprouts from the roots and bole after top-kill or damage, but the response is weak [22,86]. Most  suckers arise from suppressed buds embedded in the periderm of undisturbed roots after death or injury of aboveground parts. There is disagreement on the ability or eastern cottonwood to sprout from the bole after being cut [47,154].

Plains cottonwood does not readily form suckers or stem sprouts [87,132], and sprouting is uncommon except in flood-trained shoots [23,86,179]. 

Growth: Eastern cottonwood is the fastest growing native tree in North America [28,47,121,132,212]. It commonly increases 0.7 to 1 inch  (1.69-2.54 cm) in diameter and 5 feet (1.5 m) in height annually up to 10 to 15 years of age, and grows at only a slightly slower rate up to 30 to 35 years of age [28]. In the Mississippi valley, eastern cottonwood can reach 20 inches (50.8 cm) in diameter at breast height and 120 feet (36.6 m) in height at 35 years [28,145].

Growth of young seedlings is rapid [132]. The first 3 weeks of development of seedlings are slow, after which the growth rate rapidly accelerates [145]. Root growth of seedlings is also rapid. Seedlings have been observed to extend taproots 12 to 16 inches (30.5-40.6 cm) and lateral roots 24 inches (61 cm) by the end of their first growing season [222]. These roots serve as anchorage during floods and help to ensure a supply of water during dry periods.

Once established, seedlings are susceptible to damage from flooding. In one instance, 8 days of complete inundation weakened all seedlings, while 16 days resulted in their mortality [112]. Seedlings can survive if flooded less than 50% of the growing season [158]. Inundation does not necessarily have a detrimental effect upon germination [108].

Eastern cottonwood primarily grows on the moist alluvial soil of floodplains and bottomlands [132,145,158,171,229]. It is also found in ravines [16,107,229], along disturbed streams [117], and in low spots of sandy uplands with a high water table [229]. It also found on batture lands, the unprotected areas between the Mississippi River and its levees [145].

Plains cottonwood is found on floodplains and small sandbars in the river beds or large bends where stream flow is dramatically retarded during high water. It is found next to springs that flow long enough to form ponds [4,130,173]. 

Across its range, eastern cottonwood is found from 255 to 6,500 feet (78-1,981 m) [10,12,37,165,169,187], usually 15 to 40 feet (5-12 m) above stream level [90,151,187]. Elevation ranges for plains and Rio Grande cottonwood are given in the following table:
Plains cottonwood References
CO 3,500 to 6,500 feet (1,067-1,981 m) [49,163,180]
MT 3,200 to 4,921 feet (975-1,500 m) [49,208]
NM 5,000 to 6,500 feet (1,524-1,981 m) [147]
UT 4,500 feet (1,372 m) [49]
WY 3,500 to 9,000 feet (1,067-2,743 m) [49,69,162]
Rio Grande cottonwood
AZ 4,000 to 6,500 feet (1,219-1,981 m) [56]
CO 4,000 to 7,000 feet (1,219-2,134 m) [49]
NM 1,457 to 6,500 feet (444-1,981 m) [113,147]
TX 2,800 to 5,000 feet (853-1,524 m) [49]

The average annual precipitation across the eastern cottonwood range is 13.78 to 55 inches (350-1,397 mm) [125,169]. Average annual precipitation ranges for some states where eastern cottonwood occurs is listed below:

IN 35 to 47 inches (889-1,193.8 mm) [141]
MI 33.86 inches (860 mm) [71]
OH 34 inches (863.6 mm) [18]
TX 14.96 to 33.07 inches (380-840 mm) [213,214]
ON 37 inches (939.8 mm) [234]

Eastern cottonwood occurs in a wide range of temperature regimes with extremes ranging from -49 degrees Fahrenheit (-45 oC) in winter to 114.8 degrees Fahrenheit (46 oC) in summer [125]. The mean annual temperature or annual temperature range for some states is as follows:
MI 48.2 degrees Fahrenheit (9 oC) [71]
OH 14.5 to 83.8 degrees Fahrenheit (-9.72-28.77 oC) [18]
TX 59.9 to 67 degrees Fahrenheit (15.5-19.44 oC) [213,214]
ON 46.4 degrees Fahrenheit (8 oC) [234]

The annual precipitation varies between 12 and 30 inches (305-762 mm), averaging less than 20 inches (508 mm) across most of the plains cottonwood range. The temperature can range between -50 degrees Fahrenheit (-45.56 oC) in January and 115 degrees Fahrenheit in July (46.11 oC) [173]. The average annual precipitation and temperature for some states in the plains cottonwood range are given in the table below [4,11,15,20,36,73,82,104,105,124,127,130,146,162,175,182,186,198,208,221,231]:
IN 35 to 47 inches (889-1,193.8 mm) [141]
MI 33.86 inches (860 mm) [71]
OH 34 inches (863.6 mm) [18]
TX 14.96 to 33.07 inches (380-840 mm) [213,214]
ON 37 inches (939.8 mm) [234]

Eastern cottonwood tolerates a wide range of soils ranging from coarse sands to clays but grows best on moist, well-drained fine sandy loams or silt [47,90,129,145,151,154,158].

Plains cottonwood is found on sandy soils [124,130] or well-drained soils with a high water table to supply year-round moisture [4]. Availability of moisture is reportedly more significant to plains cottonwood than soil texture or fertility [173].

Eastern cottonwood is shade intolerant [9,47,151,171,217,225] and a pioneer species that typically establishes on freshly exposed alluvium of sandbars, streambanks, and other floodplain sites [16,23,103,117,124,150,222]. Establishment and dominance may also occur after sandbar willows have stabilized the site [229]. Eastern cottonwood invades unburned prairies in Kansas [1] and old fields and upland sites in the lake states [47,111].

Maintenance of seral, eastern cottonwood-dominated communities depends on either periodic flooding or timber harvest [23,92,155,158]. In the Great Plains, dams and reservoirs which alter the magnitude and frequency of floods, sedimentation rates, and stream meander migration rates have detrimentally affected cottonwood communities by reducing seedling recruitment and survival [23]. In the absence of flooding, succession proceeds, and the shade-intolerant cottonwoods are eventually replaced by more shade-tolerant species [155]. Eastern cottonwood stands are replaced by American elm, sycamore, pecan, sugarberry, boxelder, and sweetgum [103]. The eastern cottonwood/Rocky Mountain juniper community type is an early successional stage in North Dakota. It succeeds to the green ash/western snowberry community if left undisturbed [83].

Plains cottonwood with willow and boxelder represents an early stage on floodplains in Nebraska [4].  The plains cottonwood-willow type progresses to the plains cottonwood-green ash type in North Dakota [124]. It contributes significantly to dune formation on Lake Michigan [38].

Time of flowering and leaf emergence for eastern cottonwood varies according to geographic location. Eastern cottonwood flowers during early to late spring, generally from February through May [22,47,68,147,228] and 1 to 2 weeks before leaf initiation [22]. Seed dispersal occurs in early summer [47].  Seed dispersal and germination generally occur in late spring to early- or mid-summer and typically coincide with decreasing flow levels (May through August) [9,22,40,66,181]. Phenological dates for some states and the Great Plains are given below:

Location Flowering References Fruiting References Seed dispersal References
Eastern cottonwood            
AR March-May [115] May-June [115] ---- ----
GA April-May [230] ---- ---- ---- ----
MS ---- ---- May-August [122] ---- ----
NE April [129] ---- ---- ---- ----
OK March [222] May-June [222] May-June [222]
WV April-May [203] ---- ---- ---- ----
Plains cottonwood            
Great Plains March-June [88,147,199] May-July [88,199] ---- ----


SPECIES: Populus deltoides

Fire adaptations: Eastern cottonwood is a weak sprouter, and fire generally kills it [3,22,86,87,132,145,158]. It may sprout from the roots, root crown, or bole after fire [47,95,152,183], but sprouts are few and usually die. In a survey of postfire response of cottonwoods (Populus spp.) in Alberta, Gom and Rood [86] reported a "very poor" sprouting response for eastern cottonwood: few sprouts were produced, and most of those sprouts died. In their own study, conducted 5 months after "high-intensity" April wildfires on 2 Oldman River sites near Lethbridge, Alberta, 20% of damaged eastern cottonwood trunks produced sprouts, while 80% of damaged cottonwoods in the taxonomic section Tacamahaca produced sprouts. Five years after fire, only 10% of eastern cottonwood trunks damaged by fire still supported live sprouts.

Fire regimes: In the Northern Great Plains, historic fire frequency was influenced by topography. Where plains cottonwoods occur along rivers, the fire frequency is estimated between 20 to 30 years [189]. These riparian areas burned less frequently than the surrounding uplands; fires skip over or only burn a portion [184]. Fires most likely occurred late in the growing season when the understory vegetation was cured enough to support a fire. In the mesic portions of the Northern Great Plains where eastern cottonwood occurs, the average fire return interval is 1 to 5 years [189].

In the southern United States, "serious" fire seasons occur every 5 to 8 years. The fire season is usually in the fall, except in years with a dry, early spring [171].

Fire regimes for plant communities and ecosystems in which eastern, plains, and Rio Grande cottonwood occur are summarized below. For further information regarding fire regimes and fire ecology of these communities and ecosystems, see the 'Fire Ecology and Adaptations' section of the FEIS species summary for the plant community or ecosystem dominants listed below.

Community or Ecosystem Dominant Species Fire Return Interval Range (years)
silver maple-American elm Acer saccharinum-Ulmus americana < 35 to 200 [218]
bluestem prairie Andropogon gerardii var. gerardii-Schizachyrium scoparium < 10 [135,164]
Nebraska sandhills prairie Andropogon gerardii var. paucipilus-Schizachyrium scoparium < 10
sagebrush steppe Artemisia tridentata/Pseudoroegneria spicata 20-70 [164]
Wyoming big sagebrush Artemisia tridentata var. wyomingensis 10-70 (40**) [215,233]
plains grasslands Bouteloua spp. < 35
blue grama-buffalo grass Bouteloua gracilis-Buchloe dactyloides < 35
cheatgrass Bromus tectorum < 10 [164]
sugarberry-America elm-green ash Celtis laevigata-Ulmus americana-Fraxinus pennsylvanica < 35 to 200 [218]
northern cordgrass prairie Distichlis spicata-Spartina spp. 1-3 [164]
beech-sugar maple Fagus spp.-Acer saccharum > 1000
black ash Fraxinus nigra < 35 to 200 [218]
Rocky Mountain juniper Juniperus scopulorum < 35
wheatgrass plains grasslands Pascopyrum smithii < 35
pinyon-juniper Pinus-Juniperus spp. < 35 [164]
longleaf-slash pine Pinus palustris-P. elliottii 1-4 [159,218]
loblolly-shortleaf pine Pinus taeda-P. echinata 10 to < 35
sycamore-sweetgum-American elm Platanus occidentalis-Liquidambar styraciflua-Ulmus americana < 35 to 200 [218]
eastern cottonwood Populus deltoides 5 to 200 [164,171,189]
mesquite Prosopis glandulosa < 35 to < 100
mesquite-buffalo grass Prosopis glandulosa-Buchloe dactyloides < 35
Texas savanna Prosopis glandulosa var. glandulosa < 10 [164]
mountain grasslands Pseudoroegneria spicata 3-40 (10**) [5,6]
oak-hickory Quercus-Carya spp. < 35[218]
oak-juniper woodland (Southwest) Quercus-Juniperus spp. < 35 to < 200 [164]
white oak-black oak-northern red oak Quercus alba-Q. velutina-Q. rubra < 35
bur oak Quercus macrocarpa < 10
oak savanna Quercus macrocarpa/Andropogon gerardii-Schizachyrium scoparium 2-14 [164,218]
chestnut oak Q. prinus 3-8
northern red oak Quercus rubra 10 to < 35
black oak Quercus velutina < 35 [218]
elm-ash-cottonwood Ulmus-Fraxinus-Populus spp. < 35 to 200 [52,218]
*fire return interval varies widely; trends in variation are noted in the species summary

Tree with adventitious bud/root crown/soboliferous species root sucker
Initial off-site colonizer (off-site, initial community)


SPECIES: Populus deltoides

Fire generally kills eastern cottonwood [3,22,86,87,132,145,158]. Mature trees with thick bark may be only scarred or top-killed [145,158]. Fire scars may facilitate onset of heartwood decay [28,145,158].

No entry

Top-killed eastern cottonwood may sprout from the roots, root crown, and/or bole following fire [183,184,190]. Although studies are few, sprouting response appears to be weak, and the long-term survivability of sprouts is poor [86]. Sprouting ability of eastern cottonwood declines with age [173]. In general, the ability of cottonwoods to sprout depends on (1) species (with those in section Tacamahaca sprouting more vigorously than those in section Aigeiros), (2) age (younger trees are more efficient), and (3) location of the water table (higher water tables increase sprout survivability) [86,95].

Plains cottonwood is a weak sprouter, and is especially susceptible to late summer and fall burns [95]. It does not readily form suckers [87,132]: Root and shoot sprouts are uncommon except for flood-trained shoot suckering [179].

In an Oklahoma study, all eastern cottonwood seedlings and saplings were eliminated following a fire in June. One year later no regeneration was observed [3].

The Research Paper by Bowles and others 2007 provides information on postfire responses of several plant species, including eastern cottonwood, that was not available when this species review was written.

Prescribed fire is not recommended on bottomland forest sites in the north-central states where wood production is a primary management objective. Seedlings and young trees are easily killed by surface fires, while mature trees are often wounded. Wounded trees often contract heartrot, leading to substantial cull and volume loss [28]. Prescribed burning may stimulate sprouting and improve food supplies for wildlife species [183]. For optimum timber production, plantations must be completely protected from fire and grazing [28]. Fire prevention measures suggested for eastern cottonwood plantations include natural firebreaks (roads, trails, and sloughs) dividing stands into 40-acre (16 ha) blocks and plowed lines, at least 15 feet (4.6 m) wide [145]. To promote eastern cottonwood recovery, livestock should be excluded from burned areas for 5 years following fire and wildlife browsing should be monitored [95].

Populus deltoides: References

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