Index of Species Information

SPECIES:  Celtis reticulata

Introductory

SPECIES: Celtis reticulata
AUTHORSHIP AND CITATION : Tirmenstein, D. A. 1990. Celtis reticulata. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/ [].
ABBREVIATION : CELRET SYNONYMS : Celtis rugulosa Rydb. Celtis laevigata var. brevipes (Wats.) Sarg. Celtis laevigata Willd. var. reticulata (Torr.) [83] SCS PLANT CODE : CERE2 COMMON NAMES : netleaf hackberry western hackberry hackberry sugarberry palo blanco TAXONOMY : The scientific name of netleaf hackberry is Celtis reticulata Torr. [82]. Netleaf hackberry is a highly variable and taxonomically confusing species [41,63]. Hybridization is common within the genus Celtis and most species are poorly defined [23]. Intergrading forms and ecotypic variants are common [23]. Some authorities consider netleaf hackberry to be a variant of sugarberry (Celtis laevigata) and still others place it in synonymy with Douglas hackberry (C. douglasii) [63,73]. However, many taxonomists now regard it as a discrete species [73]. Netleaf hackberry readily hybridizes with sugarberry and populations with intermediate characteristics have been reported [74]. Some authorities delineate two forms of netleaf hackberry on the basis of leaf size [63]. Still others recognize many intergrading forms [33]. According to Kearney and others [41], "it is highly probable that more than one species is included ... as Celtis reticulata, but pending thorough revision of North American species of Celtis, no other treatment seems practicable." Biosystematic revision has been suggested for the tribe Celteae [23]. LIFE FORM : Tree, Shrub FEDERAL LEGAL STATUS : No special status OTHER STATUS : NO-ENTRY

DISTRIBUTION AND OCCURRENCE

SPECIES: Celtis reticulata
GENERAL DISTRIBUTION : Netleaf hackberry grows throughout scattered portions of the Great Basin, Pacific Northwest, and Southwest [44]. Its range extends from southern Nebraska south through central Kansas and Colorado into Texas and northern Mexico [46,63], westward to southern California, and north through Washington and Oregon into Idaho [23,46]. ECOSYSTEMS : FRES15 Oak - hickory FRES28 Western hardwoods FRES29 Sagebrush FRES30 Desert shrub FRES31 Shinnery FRES32 Texas savanna FRES33 Southwestern shrubsteppe FRES34 Chaparral - mountain shrub FRES35 Pinyon - juniper FRES40 Desert grasslands STATES : AZ CA CO ID KS NE NV NM OK OR TX UT WA MEXICO BLM PHYSIOGRAPHIC REGIONS : 2 Cascade Mountains 3 Southern Pacific Border 4 Sierra Mountains 5 Columbia Plateau 6 Upper Basin and Range 7 Lower Basin and Range 8 Northern Rocky Mountains 11 Southern Rocky Mountains 12 Colorado Plateau 13 Rocky Mountain Piedmont 14 Great Plains KUCHLER PLANT ASSOCIATIONS : K023 Juniper - pinyon woodlands K027 Mesquite bosques K031 Oak - juniper woodland K033 Chaparral K038 Great Basin sagebrush K041 Creosotebush K042 Creosotebush - bursage K044 Creosotebush - tarbush K059 Trans-Pecos shrub savanna K071 Shinnery K084 Cross Timbers K098 Northern floodplain forest SAF COVER TYPES : 63 Cottonwood 67 Mohrs ("shin") oak 68 Mesquite 239 Pinyon - juniper 240 Arizona cypress 241 Western live oak 242 Mesquite SRM (RANGELAND) COVER TYPES : NO-ENTRY HABITAT TYPES AND PLANT COMMUNITIES : Netleaf hackberry grows as an overstory dominant or codominant in a number of communities, including riparian woodlands of the Southwest and narrow gallery forests of eastern Washington and west-central Idaho. Common codominants include live oak (Quercus virginiana), cedar elm (Ulmus crassifolia), cheatgrass (Bromus tectorum), bluebunch wheatgrass (Pseudoroegneria spicata), and sand dropseed (Sporobolus cryptandrus). Published classifications listing netleaf hackberry as a dominant or indicator in community types (cts), habitat types (hts), or plant associations (pas) are presented below. Area Classification Authority w-c ID grassland and shrubland Tisdale 1986b hts, cts w-c ID, e WA riparian cts Miller and Johnson 1986 OK western oak cts Dooley and Collins 1984 OR, ID: Wallowa general veg. pas Johnson and Simon 1987 e WA, n ID steppe hts Daubenmire 1970a

MANAGEMENT CONSIDERATIONS

SPECIES: Celtis reticulata
WOOD PRODUCTS VALUE : Wood of netleaf hackberry is light brown, heavy, and not easily worked [37,43]. It is used to make boxes, crates, barrels, furniture, cabinets, paneling, and miscellaneous items [23,55,65], and is used locally for fenceposts and firewood [37,72]. Early homesteaders crafted rough furniture from hackberry wood [44]. IMPORTANCE TO LIVESTOCK AND WILDLIFE : Netleaf hackberry is used extensively for food and cover by many birds and mammals [37]. Browse: On the Edwards Plateau of Texas, netleaf hackberry is a preferred white-tailed deer browse [3,15]. In parts of southern Texas, it is a major component of mule deer diets but is relatively unimportant to white-tailed deer [2]. It may be heavily utilized by deer during drought years [1] and in southern Texas receives most use during winter and spring [15]. Pronghorn commonly browse netleaf hackberry in the spring [11]. Beaver feed on hackberry wood in many areas [48]. Scrub jays commonly feed on leaf galls present on foliage of netleaf hackberry [34]. Cattle sometimes browse netleaf hackberry [59], although it is most often used on overgrazed sites where more preferred forage is unavailable [20]. Spanish goats often seek out tender young sprouts during the first year after fire [70]. Fruit: Fruit of netleaf hackberry is readily consumed by many birds [55,73]. It is considered the single most important winter bird food at the lower edge of the mountain brush zone along the Wasatch Front of northern Utah [34]. The band-tailed pigeon, Steller's jay, northern flicker, American robin, Townsend's solitaire, Bohemian waxwing, cedar waxwing, American crow, scrub jay, and rufous-sided towhee feed on this persistent berry [34,43]. It reportedly constitutes an emergency food source for avian seed eaters during January and February [62]. Many mammals, including squirrels, foxes, Barbary sheep, and coyotes, also eat netleaf hackberry fruit [43,76]. PALATABILITY : Leaves of netleaf hackberry become somewhat tough as they mature [74] and may decline in palatability to some species; however, white-tailed deer preference for this species tends to be highest in summer and fall [60,62]. In Texas, general palatability has been rated as follows [11]: Pronghorn excellent Cattle poor Domestic sheep poor Fruit of netleaf hackberry is highly palatable to many birds and mammals [48]. NUTRITIONAL VALUE : Netleaf hackberry browse in the Edwards Plateau of Texas has been rated as good in protein (14.35%), good in phosphoric acid (P2O5) (0.38%), and fair in lime (CaO) (6.27%) [26]. COVER VALUE : Netleaf hackberry provides good cover for a variety of big game species [39,43]. The dense cover of netleaf hackberry stands is favored by white-tailed deer in the Rio Grande Valley of southern Texas [49]. Southern plains woodrats use netleaf hackberry twigs to construct houses [66]. Netleaf hackberry provides nesting sites for the white-tailed raven, Swainson's hawk, scissor-tailed flycatcher, Bullock's oriole, and many doves, quail, and numerous desert songbirds [19,27,43]. Hackberries offer good hiding or resting cover for quail in many parts of the Southwest [27]. Netleaf hackberry provides much sought-after shade for domestic livestock in the Southwest and in the Snake River Drainage of Idaho [18,39]. VALUE FOR REHABILITATION OF DISTURBED SITES : Netleaf hackberry can be used to aid in soil stabilization on various types of disturbed sites [54,67]. It is well adapted to mountain-brush and pinyon-juniper communities [54]. Netleaf hackberry can be propagated from seed, which when cleaned, averages 4,870 per pound (10,727/kg) [8]. It can also be propagated vegetatively from stem cuttings [8]. Nursery or container stock can be transplanted onto disturbed sites with good results [54,64]. OTHER USES AND VALUES : Netleaf hackberry is well suited for use in landscaping [38]. This small shade tree is tolerant of dry sites and can be planted in yards or patios [23,64], and along streets in urban areas [23]. The shade value of netleaf hackberry was also recognized by early Native American peoples, including the Basketmakers of the Southwest [44]. Because of its tendency to grow near flowing water, this tree provided the focus for habitations such as Hovenweep and Montezuma Castle [44]. The sweet, edible fruit [37,63] was traditionally an important food source for many Native American peoples [55,72]. Today, the fruit receives only limited human use [37]. OTHER MANAGEMENT CONSIDERATIONS : Netleaf hackberry is susceptible to "witches broom" and various insect infestations [37,44,75]. Netleaf hackberry can be reduced by heavy grazing [59].

BOTANICAL AND ECOLOGICAL CHARACTERISTICS

SPECIES: Celtis reticulata
GENERAL BOTANICAL CHARACTERISTICS : Netleaf hackberry is a spreading, scraggly, often stunted tree or large shrub [44,55,63,75]. It commonly grows from 7 to 20 feet (2-6 m) in height but can reach up to 53 feet (16 m) in height and 24 inches (60 cm) in diameter on favorable sites [31,53]. Plants generally grow slowly [64] and live for 100 to 200 years [62]. The trunk is usually short and crooked [4,44] with thick, warty, reddish-brown to gray bark [37,44]. Slender twigs are reddish-brown, glabrous or puberulent [53], and often form a twisting network [4]. Plants are strongly taprooted but possess many shallow roots as well [62]. Roots are fibrous, wide spreading, and can reach maximum depths of 15 feet (5 m) or more [64,80]. RAUNKIAER LIFE FORM : Phanerophyte REGENERATION PROCESSES : Flowering and fruiting: Netleaf hackberry is monoecious [63]. Small, inconspicuous green flowers are borne singly in the leaf axils on current year's twigs [33,37,63]. The fruit is an orange to reddish, purplish, or black drupe that contains a single seed or nutlet [33]. The fruit is sweet with thin, dry pulp [4,76]. Seed: Netleaf hackberry produces an abundance of persistent seed nearly every year [8]. The hard, bony seeds are cream-colored and approximately 0.22 inch (5.5 mm) in diameter [4,63]. Seed is readily dispersed by a variety of birds and mammals [44]. Seed remains viable under laboratory conditions for at least several years [8]; seed longevity under natural conditions has not been documented. Germination: Seed dormancy can be broken by stratification for 120 days at 41 degrees F (5 degrees C) [8]. Germination is also enhanced by depulping the fruit prior to planting [8]. Germination has averaged 37 to 80 percent in laboratory tests [8,72]. Natural germination occurs in late winter and spring [80]. Seedling establishment: Seedlings are commonly observed on moist loamy drainageway soils, in the sand of ephemeral streambeds, and in saturated alluvium in waterways with sustained flows [22,80]. Establishment may be favored on high terraces in riparian zones where floods may disturb the channels themselves but leave terraces relatively undisturbed [5]. Vegetative regeneration: Netleaf hackberry sprouts from the root crown after aboveground portions of the plant are removed or damaged [14]. SITE CHARACTERISTICS : Netleaf hackberry commonly grows in bottomlands, washes, ravines, arroyos, rocky canyons, and along streamcourses, water tanks, and ponds [31,33,37,55,79]. Netleaf hackberry is particularly abundant in floodplain forests along large rivers of the Edwards Plateau of Texas [71,79] and is common in gallery forests along the major canyons of Snake, Salmon, and Columbia River valleys [18,50,68]. Netleaf hackberry also occurs as scattered individuals in desert shrubland and in semidesert grassland communities [9,28]. Scattered individuals often occur where upper desert grassland communities grade into low savannas [21]. In Texas, netleaf hackberry is particularly common on rolling plains and breaks, and as a component of hill and bluff timber [25,55,62]. Netleaf hackberry occurs in Great Basin montane scrubland, creosotebush scrub, and wash scrub communities, pinyon-juniper and Joshua tree woodlands, and mesquite bosques of the Southwest [10,13,30,52,53]. Plant associates: Common associates in Southwestern riparian woodland communities include walnut (Juglans spp.), willow (Salix spp.), cottonwood (Populus spp), American sycamore (Platanus occidentalis), Texas mulberry (Morus microphylla), western soapberry (Sapindus saponaria var. drummondii), live oak (Quercus virginana), cedar elm (Ulmus crassifolia), ash (Fraxinus spp.), Texas persimmon (Diospyros texana), and mesquite (Prosopis glandulosa) [52,78,79]. Overstory codominants in terrace communities include live oak, little walnut (Juglans microcarpa), and pecan (Carya illinoensis) [79]. Other common Southwestern associates include Utah juniper (Juniperus osteosperma), Ashe juniper (J. ashei), live oak (Quercus fusiformis), black cherry (Prunus serotina), whitethorn acacia (Acacia constricta), fendlerbush (Fallugia paradoxa) [12,13,81]. Cheatgrass, white alder (Alnus rhombifolia), water birch (Betula occidentalis), black poplar (Populus trichocarpa), sand dropseed, bluebunch wheatgrass, and Kentucky bluegrass (Poa pratensis) occur with netleaf hackberry in gallery forests of Idaho and Washington [18,50,68]. Soils: Netleaf hackberry grows on well-drained, dry to moist soils [64]. It occurs on gravelly or rocky soils, and also on sand and loam [22,31,72]. Netleaf hackberry grows on alkaline or acidic soils [54], but pH averages 7.0 to 7.5 [64]. Soils are commonly derived from limestone, or alluvial or colluvial parent materials [31,39,68]. Near the Gulf, plants occur on calcareous shell banks [73]. In many areas, netleaf hackberry develop best on alluvial soils [71]. Climate: Netleaf hackberry grows well in sun and is tolerant of drought [38,72]. It occurs in subhumid to semiarid areas characterized by mesothermal climatic regimes [71]. Average annual precipitation ranges from 15 to 33 inches (38-84 cm) in parts of Texas [71]; however, netleaf hackberry can grow where annual precipitation averages only 7 inches (18 cm) [4]. In some areas, summertime temperatures may exceed 110 degrees F (43 degrees C) [4]. Netleaf hackberry may be restricted by soil moisture levels [79]. Its distribution may also be largely limited by flood tolerance. Elevation: In many areas, netleaf hackberry is restricted to waterways and spans a considerable elevational range [32]. It grows at elevations ranging from 2,500 to 6,000 feet (762-1,829 m) in Arizona [41]; from 2,800 to 5,000 feet (853-1,524 m) in California [53]; and up to 6,000 feet (2,000 m) in Utah [37]. SUCCESSIONAL STATUS : Netleaf hackberry can invade many types of newly disturbed sites but can also persist in a number of climax communities where soil-water regimes are favorable. Texas: On the Edwards Plateau, netleaf hackberry is among the large-seeded facultative riparian species that invade river terrace communities [5]. It also increases after prescribed burns in Ashe juniper communities on toeslopes where water availability is relatively high [57]. Although it commonly codominates these moister toeslopes with flameleaf sumac (Rhus lanceolata) and live oak (Quercus fusiformis), it is often absent on adjacent drier sites [57]. Idaho-eastern Washington: In the middle Snake and lower Salmon drainages of west-central Idaho and eastern Washington, netleaf hackberry occurs as an overstory dominant on lower valley slopes and alluvial terraces [18,68]. Many of these sites have been disturbed by domestic livestock. Current understory dominants on disturbed sites include annual bromes (cheatgrass, Japanese brome (B. japonicus), and poverty brome (B. sterilis), and sand dropseed, but evidence suggests that bluebunch wheatgrass grew as an understory dominant beneath netleaf hackberry prior to disturbance [68]. Daubenmire [18] identified a netleaf hackberry/cheatgrass habitat type, although Tisdale [68] maintained that "designation of a climax community type with an exotic annual as one of the dominants seems inappropriate, especially when stands of Celtis douglasii [reticulata]/ bluebunch wheatgrass can be found on similar habitats in Washington along the Grande Ronde River." Tallgrass prairie: Plowing in the tallgrass prairie disturbs the roots of grasses, reducing their competitive ability and allowing for the subsequent establishment of weeds and woody invaders. Woody plants such as netleaf hackberry tend to increase after annual cultivation in tallgrass prairies of central Oklahoma [16]. Grass cover typically increases immediately after plowing stops and peaks 5 years later. Sumac (Rhus spp.), indian-currant coralberry (Symphoricarpos orbiculatus), and roughleaf dogwood (Cornus drummondii) soon become established, and netleaf hackberry and slippery elm (Ulmus rubra) gradually invade the area. Although woody plants remain rare during the first 9 years, these invaders begin to increase dramatically as grassland vegetation loses dominance. Netleaf hackberry, indian-currant coralberry, flameleaf sumac (Rhus copallina), and weedy grasses such as johnson-grass (Sorghum halepense) dominate some 9- to 32-year-old stands. Collins and Adams [16] noted that succession can be both rapid and unpredictable in the tallgrass prairie, and "even at the physiognomic level, general successional trends may be difficult to quantify." SEASONAL DEVELOPMENT : Leaves of netleaf hackberry first appear in early April to late May [23] and mature in June [74]. Plants flower in spring, with or shortly after initial leaf development [23]. Fruit ripens in late summer or fall [64]. Fruit may persist through the winter [62], although some seed is dispersed during the fall and winter [8,72]. Generalized flowering and fruiting dates by geographic location are as follows: Location Flowering Fruit ripe Authority c Great Plains late April August-Sept. Stephens 1973 s CA April-May ---- Munz 1974 Great Plains April-Sept. ---- Great Plains Flora Association 1986

FIRE ECOLOGY

SPECIES: Celtis reticulata
FIRE ECOLOGY OR ADAPTATIONS : Netleaf hackberry is often associated with riparian woodlands which burn infrequently. These narrow canyon or gallery forests contrast strikingly with adjacent desert shrublands or grasslands where netleaf hackberry occurs as scattered individuals. Recurrent fires in drier upland types may eliminate or reduce invading shrubs and trees [32]. Netleaf hackberry also persists in fire-prone toeslope communities of Texas [57]. It commonly sprouts from the stem base or root crown after fire [57,70] and becomes prominent in many postburn communities. Birds and mammals presumably transport some seed from adjacent unburned areas [44], and limited postfire seedling establishment is possible. POSTFIRE REGENERATION STRATEGY : survivor species; on-site surviving root crown or caudex off-site colonizer; seed carried by animals or water; postfire yr 1&2

FIRE EFFECTS

SPECIES: Celtis reticulata
IMMEDIATE FIRE EFFECT ON PLANT : Netleaf hackberry is described as fairly tolerant of fire [3]. Portions of the root crown commonly survive after aboveground vegetation is consumed by fire [3,57,70]. Plants are rarely killed by fire [14]. DISCUSSION AND QUALIFICATION OF FIRE EFFECT : NO-ENTRY PLANT RESPONSE TO FIRE : Netleaf hackberry sprouts from the root crown after aboveground vegetation is consumed by fire [3,14]. In some instances, recovery may be relatively rapid and cover can increase dramatically. Netleaf hackberry can reportedly outcompete species such as agarito (Mahonia trifoliolata) in early postburn communities [59]. On the Edwards Plateau, netleaf hackberry readily sprouted and increased in canopy cover after prescribed fire and mechanical scarification [57]. After fire it codominated (18 percent cover) relatively moist toeslopes with flameleaf sumac (22 percent cover), and live oak (Quercus fusiformis) (20 percent cover). Very little netleaf hackberry was observed on unburned areas or on drier burned sites (< 1 percent) [57]. DISCUSSION AND QUALIFICATION OF PLANT RESPONSE : NO-ENTRY FIRE MANAGEMENT CONSIDERATIONS : Prescribed fire: Bock and Bock [7] reported that prescribed fire is "difficult to manage and potentially very destructive" in established riparian woodlands of the Southwest. These relatively rare and fragile areas provide important food and cover for desert wildlife [61]. Because browse and cover are often limited in these areas, burning is not generally recommended [63]. Wildlife: Removal of shrub-dominated communities can adversely impact wildlife in many areas. Deer commonly avoid open areas, and if burning is planned in shrub communities, efforts should be made to burn in mosaics, leaving strips of cover [3,49]. In some instances, it may be advisable to leave drainages intact for deer use [49]. On the Edwards Plateau of Texas, burning at 7- to 10-year intervals is recommended if management aims include controlling shrubs while maintaining deer populations. Deer numbers can be reduced if burns are conducted at more frequent intervals [3]. Burning woody vegetation in some shrub-grassland communities can be detrimental to birds, especially if conducted during the breeding season [19].

REFERENCES

SPECIES: Celtis reticulata
REFERENCES : 1. Anthony, Robert G. 1976. Influence of drought on diets and numbers of desert deer. Journal of Wildlife Management. 40(1): 140-144. [11558] 2. Anthony, Robert G.; Smith, Norman S. 1977. Ecological relationships between mule deer and white-tailed deer in southeastern Arizona. Ecological Monographs. 47: 255-277. [9890] 3. Armstrong, W. E. 1980. Impact of prescribed burning on wildlife. In: White, Larry D., ed. Prescribed range burning in the Edwards Plateau of Texas: Proceedings of a symposium; 1980 October 23; Junction, TX. College Station, TX: Texas Agricultural Extension Service, The Texas A&M University System: 22-26. [11430] 4. Arno, Stephen F.; Hammerly, Ramona P. 1977. Northwest trees. Seattle, WA: The Mountaineers. 222 p. [4208] 5. Asplund, Kenneth K.; Gooch, Michael T. 1988. Geomorphology and the distributional ecology of Fremont cottonwood (Populus fremontii) in a desert riparian canyon. Desert Plants. 9(1): 17-27. [563] 6. Bernard, Stephen R.; Brown, Kenneth F. 1977. Distribution of mammals, reptiles, and amphibians by BLM physiographic regions and A.W. Kuchler's associations for the eleven western states. Tech. Note 301. Denver, CO: U.S. Department of the Interior, Bureau of Land Management. 169 p. [434] 7. Bock, Carl E.; Bock, Jane H. 1990. Effects of fire on wildlife in southwestern lowland habitats. In: Krammes, J. S., technical coordinator. Effects of fire management of Southwestern natural resources: Proceedings of the symposium; 1988 November 15-17; Tucson, AZ. Gen. Tech. Rep. RM-191. Fort Collins, CO: U.S. Department of Agriculture, Forest Service, Rocky Mountain Forest and Range Experiment Station: 50-64. [11273] 8. Bonner, F. T. 1974. Celtis L. Hackberry. In: Schopmeyer, C. S., technical coordinator. Seeds of woody plants in the United States. Agric. Handb. 450. Washington, DC: U.S. Department of Agriculture, Forest Service: 298-300. [7579] 9. Bowers, Janice E.; McLaughlin, Steven P. 1987. Flora and vegetation of the Rincon Mountains, Pima County, Arizona. Desert Plants. 8(2): 50-94. [495] 10. Brown, David E. 1982. Great Basin montane scrubland. In: Brown, David E., ed. Biotic communities of the American Southwest--United States and Mexico. Desert Plants. 4(1-4): 83-84. [8890] 11. Buechner, Helmut K. 1950. Life history, ecology, and range use of the pronghorn antelope in Trans-Pecos Texas. American Midland Naturalist. 43(2): 257-354. [4084] 12. Campbell, C. J. 1973. Pressure bomb measurements indicate water availability in a southwestern riparian community. Res. Note RM-246. Fort Collins, CO: U.S. Department of Agriculture, Forest Service, Rocky Mountain Forest and Range Experiment Station. 4 p. [11155] 13. Carignan, Jeanette M. 1988. Ecological survey and elevational gradient implications of the flora and vertebrate fauna in the northern Del Norte Mountains, Brewster Co., Tx. Alpine, TX: Sul Ross State University. 181 p. Thesis. [12255] 14. Carter, Meril G. 1. Effects of drougth on mesquite. Journal of Range Management. 17: 275-276. [10176] 15. Chamrad, Albert D.; Box, Thadis W. 1968. Food habits of white-tailed deer in south Texas. Journal of Range Management. 21: 158-164. [10857] 16. Collins, S. L.; Adams, D. E. 1983. Succession in grasslands: thirty-two years of change in a central Oklahoma tallgrass prairie. Vegetatio. 51: 181-190. [2929] 17. Cox, Mary Josephine. 1941. The comparative anatomy of the secondary xylem of five American species of Celtis. American Midland Naturalist. 25: 348-357. [11743] 18. Daubenmire, R. 1970. Steppe vegetation of Washington. Technical Bulletin 62. Pullman, WA: Washington State University, College of Agriculture, Washington Agricultural Experiment Station. 131 p. [733] 19. Davis, C. A.; Sawyer, P. E.; Griffing, J. P.; Borden, B. D. 1974. Bird populations in a shrub-grassland area, southeastern New Mexico. Bulletin 619. Las Cruces, NM: New Mexico State University, Agricultural Experiment Station. 29 p. [4548] 20. Dayton, William A. 1931. Important western browse plants. Misc. Publ. 101. Washington, DC: U.S. Department of Agriculture. 214 p. [768] 21. Dick-Peddie, W. A.; Moir, W. H. 1970. Vegetation of the Organ Mountains, New Mexico. Science Series No. 4. Fort Collins, CO: Colorado State University, Range Science Department. 28 p. [6699] 22. Dooley, Karen L.; Collins, Scott L. 1984. Ordination and classification of western oak forests in Oklahoma. American Journal of Botany. 71(9): 1221-1227. [11543] 23. Elias, Thomas S. 1970. The genera of Ulmaceae in the southeastern United States. Journal of the Arnold Arboretum. 51: 18-40. [11742] 24. Eyre, F. H., ed. 1980. Forest cover types of the United States and Canada. Washington, DC: Society of American Foresters. 148 p. [905] 25. Foster, J. H.; Krausz, H. B.; Leidigh, A. H. 1917. General survey of Texas woodlands including a study of the commercial possibilities of mesquite. Bulletin of the Agricultural and Mechanical College of Texas, Third Series. College Station, TX: Agricultural and Mechanical College of Texas, Department of Forestry; 3(9): 1-47. [11796] 26. Fraps, G. S.; Cory, V. L. 1940. Composition and utilization of range vegetation of Sutton and Edwards Counties. Bulletin No. 58. College Station, TX: Texas Agricultural Experiment Station. 39 p. [5746] 27. Fulbright, Timothy E.; Flenniken, Kay S.; Waggerman, Gary L. 1986. Enhancing germination of spiny hackberry seeds. Journal of Range Management. 39(6): 552-554. [3020] 28. Gardner, J. L. 1950. Effects of thirty years of protection from grazing in desert grassland. Ecology. 31(1): 44-50. [4423] 29. Garrison, George A.; Bjugstad, Ardell J.; Duncan, Don A.; [and others]. 1977. Vegetation and environmental features of forest and range ecosystems. Agric. Handb. 475. Washington, DC: U.S. Department of Agriculture, Forest Service. 68 p. [998] 30. Glinski, Richard L. 1977. Regeneration and distribution of sycamore and cottonwood trees along Sonoita Creek, Santa Cruz County, Arizona. In: Johnson, R. Roy; Jones, Dale A., tech. coords. Importance, preservation and management of riparian habitat: a symposium: Proceedings; 1977 July 9; Tucson, AZ. Gen. Tech. Rep. RM-43. Fort Collins, CO: U.S. Department of Agriculture, Forest Service, Rocky Mountain Forest and Range Experiment Station: 116-123. [5340] 31. Great Plains Flora Association. 1986. Flora of the Great Plains. Lawrence, KS: University Press of Kansas. 1392 p. [1603] 32. Hastings, James R.; Turner, Raymond M. 1965. The changing mile: An ecological study of vegetation change with time in the lower mile of an arid and semiarid region. Tuscon, AZ: University of Arizona Press. 317 p. [10533] 33. White, E. M. 1961. A possible relationship of little bluestem distribution to soils. Journal of Range Management. 14: 243-247. [110] 34. Hayward, C. Lynn. 1948. Biotic communities of the Wasatch Chaparral, Utah. Ecological Monographs. 18: 473-506. [9633] 35. Hitchcock, C. Leo; Cronquist, Arthur. 1973. Flora of the Pacific Northwest. Seattle, WA: University of Washington Press. 730 p. [1168] 36. Huston, J. E.; Rector, B. S.; Merrill, L. B.; Engdahl, B. S. 1981. Nutritional value of range plants in the Edwards Plateau region of Texas. Report B-1375. College Station, TX: Texas A&M University System, Texas Agricultural Experiment Station. 16 p. [4565] 37. Johnson, Carl M. 1970. Common native trees of Utah. Special Report 22. Logan, UT: Utah State University, College of Natural Resources, Agricultural Experiment Station. 109 p. [9785] 38. Gutknecht, Kurt W. 1989. Xeriscaping: an alternative to thirsty landscapes. Utah Science. 50(4): 142-146. [10166] 39. Johnson, Charles G., Jr.; Simon, Steven A. 1987. Plant associations of the Wallowa-Snake Province: Wallowa-Whitman National Forest. R6-ECOL-TP-255A-86. Baker, OR: U.S. Department of Agriculture, Forest Service, Pacific Northwest Region, Wallowa-Whitman National Forest. 399 p. [9600] 40. Kartesz, John T.; Kartesz, Rosemarie. 1980. A synonymized checklist of the vascular flora of the United States, Canada, and Greenland. Volume II: The biota of North America. Chapel Hill, NC: The University of North Carolina Press; in confederation with Anne H. Lindsey and C. Richie Bell, North Carolina Botanical Garden. 500 p. [6954] 41. Kearney, Thomas H.; Peebles, Robert H.; Howell, John Thomas; McClintock, Elizabeth. 1960. Arizona flora. 2d ed. Berkeley, CA: University of California Press. 1085 p. [6563] 42. Kuchler, A. W. 1964. Manual to accompany the map of potential vegetation of the conterminous United States. Special Publication No. 36. New York: American Geographical Society. 77 p. [1384] 43. Lamb, S. H. 1971. Woody plants of New Mexico and their value to wildlife. Bull. 14. Albuquerque, NM: New Mexico Department of Game and Fish. 80 p. [9818] 44. Lanner, Ronald M. 1983. Trees of the Great Basin: A natural history. Reno, NV: University of Nevada Press. 215 p. [1401] 45. Little, Elbert L., Jr. 1976. Atlas of United States trees. Volume 3. Minor western hardwoods. Misc. Publ. 1314. Washington, DC: U.S. Department of Agriculture, Forest Service. 13 p. 290 maps. [10430] 46. Little, Elbert L., Jr. 1979. Checklist of United States trees (native and naturalized). Agric. Handb. 541. Washington, DC: U.S. Department of Agriculture, Forest Service. 375 p. [2952] 47. Lyon, L. Jack; Stickney, Peter F. 1976. Early vegetal succession following large northern Rocky Mountain wildfires. In: Proceedings, Tall Timbers fire ecology conference and Intermountain Fire Research Council fire and land management symposium; 1974 October 8-10; Missoula, MT. No. 14. Tallahassee, FL: Tall Timbers Research Station: 355-373. [1496] 48. Martin, Alexander C.; Zim, Herbert S.; Nelson, Arnold L. 1951. American wildlife and plants. New York: McGraw-Hill Book Company, Inc. 500 p. [4021] 49. McMahan, Craig A.; Inglis, Jack. 1974. Use of Rio Grande plain brush types by white-tailed deer. Journal of Range Management. 27(5): 369-374. [11557] 50. Miller, Thomas B.; Johnson, Frederic D. 1986. Sampling and data analyses of narrow, variable-width gallery forests over environmental gradients. Tropical Ecology. 27: 132-142. [12310] 51. Minckley, W. L.; Brown, David E. 1982. Wetlands. In: Brown, David E., ed. Biotic communities of the American Southwest--United States and Mexico. Desert Plants. 4(1-4): 223-287. [8898] 52. Minckley, W. L.; Clark, Thomas O. 1981. Vegetation of the Gila River Resource Area, eastern Arizona. Desert Plants. 3(3): 124-140. [10863] 53. Munz, Philip A. 1974. A flora of southern California. Berkeley, CA: University of California Press. 1086 p. [4924] 54. Plummer, A. Perry. 1977. Revegetation of disturbed Intermountain area sites. In: Thames, J. C., ed. Reclamation and use of disturbed lands of the Southwest. Tucson, AZ: University of Arizona Press: 302-337. [171] 55. Powell, A. Michael. 1988. Trees & shrubs of Trans-Pecos Texas including Big Bend and Guadalupe Mountains National Parks. Big Bend National Park, TX: Big Bend Natural History Association. 536 p. [6130] 56. Preston, Richard J., Jr. 1948. North American trees. Ames, IA: The Iowa State College Press. 371 p. [1913] 57. Rasmussen, G. Allen; Wright, Henry A. 1989. Succession of secondary shrubs on Ashe juniper communities after dozing and prescribed burning. Journal of Range Management. 42(4): 295-298. [7856] 58. Raunkiaer, C. 1934. The life forms of plants and statistical plant geography. Oxford: Clarendon Press. 632 p. [2843] 59. Schmidt, Harold. 1980. Improving shinoak range with prescribed fire. In: White, Larry D., ed. Prescribed range burning in the Edwards Plateau of Texas: Proceedings of a symposium; 1980 October 23; Junction, TX. College Station, TX: Texas Agricultural Extension Service, The Texas A&M University System: 45-47. [11432] 60. Severson, Kieth E.; Medina, Alvin L. 1983. Deer and elk habitat management in the Southwest. Journal of Range Management Monograph No. 2. Denver: Society for Range Management. 64 p. [2110] 61. Severson, Kieth E.; Rinne, John N. 1990. Increasing habitat diversity in Southwestern forests and woodlands via prescribed fire. In: Krammes, J. S., technical coordinator. Effects of fire management of Southwestern natural resources: Proceedings of the symposium; 1988 November 15-17; Tucson, AZ. Gen. Tech. Rep. RM-191. Fort Collins, CO: U.S. Department of Agriculture, Forest Service, Rocky Mountain Forest and Range Experiment Station: 94-104. [11277] 62. Simpson, Benny J. 1988. A field guide to Texas trees. Austin, TX: Texas Monthly Press. 372 p. [11708] 63. Stephens, H. A. 1973. Woody plants of the North Central Plains. Lawrence, KS: The University Press of Kansas. 530 p. [3804] 64. Sutton, Richard F.; Johnson, Craig W. 1974. Landscape plants from Utah's mountains. EC-368. Logan, UT: Utah State University, Cooperative Extension Service. 135 p. [49] 65. Sweitzer, Edward M. 1971. Comparative anatomy of Ulmaceae. Journal of the Arnold Arboretum. 52(4): 523-585. [11744] 66. Thies, Monte; Caire, William. 1990. Association of Neotoma micropus nests with various plant species in southwestern Oklahoma. Southwestern Naturalist. 35(1): 80-102. [11140] 67. Thornburg, Ashley A. 1982. Plant materials for use on surface-mined lands. SCS-TP-157. Washington, DC: U.S. Department of Agriculture, Soil Conservation Service. 88 p. [3769] 68. Tisdale, E. W. 1986. Canyon grasslands and associated shrublands of West-central Idaho and adjacent areas. Bulletin Number 40. Moscow, ID: University of Idaho, Forest, Wildlife and Range Experiment Station, College of Forestry, Wildlife and Range Sciences. 42 p. [2338] 69. U.S. Department of Agriculture, Soil Conservation Service. 1982. National list of scientific plant names. Vol. 1. List of plant names. SCS-TP-159. Washington, DC. 416 p. [11573] 70. Ueckert, Darrell N. 1980. Manipulating range vegetation with prescribed fire. In: White, Larry D., ed. Prescribed range burning in the Edwards Plateau of Texas: Proceedings of a symposium; 1980 October 23; Junction, TX. College Station, TX: Texas Agricultural Extension Service, The Texas A&M University System: 27-44. [11431] 71. Van Auken, O. W.; Ford, A. L.; Stein, A. 1979. A comparison of some woody upland and riparian plant communities of the southern Edwards Plateau. Southwestern Naturalist. 24(1): 165-180. [10489] 72. Van Dersal, William R. 1938. Native woody plants of the United States, their erosion-control and wildlife values. Washington, DC: U.S. Department of Agriculture. 362 p. [4240] 73. Vines, Robert A. 1960. Trees, shrubs, and woody vines of the Southwest. Austin, TX: University of Texas Press. 1104 p. [7707] 74. Waller, D. A. 1982. Leaf-cutting ants and avoided plants: defenses against Atta texana attack. Oecologia. 52(3): 400-403. [12160] 75. Weber, William A. 1987. Colorado flora: western slope. Boulder, CO: Colorado Associated University Press. 530 p. [7706] 76. Welsh, Stanley L.; Atwood, N. Duane; Goodrich, Sherel; Higgins, Larry C., eds. 1987. A Utah flora. Great Basin Naturalist Memoir No. 9. Provo, UT: Brigham Young University. 894 p. [2944] 77. Wheeler, E. A.; LaPasha, C. A.; Miller, R. B. 1989. Wood anatomy of elm (Ulmus) and hackberry (Celtis) species native to the United States. International Association of Wood Anatomy Bulletin. 10(1): 5-26. [11552] 78. Wood, Carl E.; Wood, Judith K. 1988. Woody vegetation of the Frio River riparian forest, Texas. Texas Journal of Science. 40(3): 309-322. [11870] 79. Wood, Carl E.; Wood, Judith K. 1989. Riparian forests of the Leona and Sabinal Rivers. Texas Journal of Science. 41(4): 395-412. [11869] 80. Zimmermann, Robert C. 1969. Plant ecology of an arid basin: Tres Alamos-Redington Area, southeastern Arizona. Geological Survey Professional Paper 485-D. Washington, DC: U.S. Department of the Interior, Geological Survey. 51 p. [4287] 81. Van Auken, O. W.; Ford, A. L.; Stein, A.; Stein, A. G. 1980. Woody vegetation of upland plant communities in the southern Edwards Plateau. Texas Journal of Science. 32: 23-35. [10859] 82. Hickman, James C., ed. 1993. The Jepson manual: Higher plants of California. Berkeley, CA: University of California Press. 1400 p. [21992] 83. Kartesz, John T.; Meacham, Christopher A. 1999. Synthesis of the North American flora (Windows Version 1.0), [CD-ROM]. Available: North Carolina Botanical Garden. In cooperation with the Nature Conservancy, Natural Resources Conservation Service, and U.S. Fish and Wildlife Service [2001, January 16]. [36715]


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