Index of Species Information
SPECIES: Salix richardsonii
SPECIES: Salix richardsonii
AUTHORSHIP AND CITATION :
Esser, Lora L. 1992. Salix richardsonii.
In: Fire Effects Information System, [Online].
U.S. Department of Agriculture, Forest Service,
Rocky Mountain Research Station, Fire Sciences Laboratory (Producer).
Available: http://www.fs.fed.us/database/feis/ .
SCS PLANT CODE :
COMMON NAMES :
The currently accepted scientific name for Richardson willow is Salix
richardsonii Hook. . There are no varieties, forms, or natural hybrids;
although hybridization and introgression with S. barclayi and with S.
alaxensis var. alaxensis has been suggested .
LIFE FORM :
FEDERAL LEGAL STATUS :
No special status
OTHER STATUS :
DISTRIBUTION AND OCCURRENCE
SPECIES: Salix richardsonii
GENERAL DISTRIBUTION :
Richardson willow occurs from the Arctic Coast southward through most of
central and south-central Alaska; it does not occur in the western
Alaskan Peninsula or Kenai Peninsula. It extends eastward across
northern Canada to the Baffin Islands; southward to northwest Hudson
Bay; and west to northern British Columbia [1,5,19,35].
AK BC NT YT
BLM PHYSIOGRAPHIC REGIONS :
KUCHLER PLANT ASSOCIATIONS :
K052 Alpine meadows and barren
SAF COVER TYPES :
201 White spruce
202 White spruce - paper birch
203 Balsam poplar
204 Black spruce
251 White spruce - aspen
252 Paper birch
253 Black spruce - white spruce
254 Black spruce - paper birch
SRM (RANGELAND) COVER TYPES :
HABITAT TYPES AND PLANT COMMUNITIES :
Richardson willow is a common, thicket-forming shrub of streambanks and
moist slopes in the Arctic and above timberline where it is often
associated with alders (Alnus spp.) and birch (Betula spp.); it is also
found in open spruce (Picea spp.) stands and old burns at lower
elevations [5,35]. Richardson willow can also be found in floodplain
thickets on rivers and grows on newly exposed alluvial deposits that are
periodically flooded [12,35]. Individual shrubs can be found on
south-facing steppes, on pingos found in permafrost regions, and on dry,
rock outcrops [13,39]. Published classifications describing Richardson
willow as a codominant in community types are listed below:
Arctic community types of Northwest Alaska .
SPECIES: Salix richardsonii
IMPORTANCE TO LIVESTOCK AND WILDLIFE :
Richardson willow is an important food source for moose, caribou, mule
deer, muskrat, and beaver [13,23,24,26,31]. During the winter in
Alaska, moose feed primarily on shoots of current growth of willow
(Salix spp.), quaking aspen (Populus tremuloides), paper birch (Betula
papyrifera), and balsam poplar (Populus balsamifera) growing as shrubs
or saplings in young, seral communities. Of the preceding species,
willow is the most preferred by moose . Of the more than 20 species
of willow found in Denali National Park, Richardson willow was one of
three species utilized the most . In one study, willows comprised
94 percent of total biomass consumed by moose from January to April,
with Richardson willow comprising 6.1 percent .
Richardson willow is considered moderately palatable. In one study,
Richardson willow was consumed by moose to a greater extent when
occurring in mixed stands with highly preferred species than when
growing in pure stands . Willow palatability increases as the
season progresses .
NUTRITIONAL VALUE :
Nutrient composition of Richardson willow consumed by moose in Denali
National Park, Alaska, from January to April, 1984, was as follows :
gross energy: 5.08 kcal/g
percent in vitro digestible organic matter: 39.4
percent of dry matter: 5.9
percent lignin: 19.4
percent ash: 2.8
percent ether extract: 8.3
The spring protein concentration of willow twigs is three times greater
than that of willow bark. Calcium concentration is greater in bark than
in twigs, and phosphorus concentration is greater in twigs than in bark
COVER VALUE :
Richardson willow characteristically produces dense thickets along
streams and rivers, which provide thermal and hiding cover for mule
deer. Branches are used by beaver in the construction of dams and lodges .
Richardson willow can stabilize streambanks when thickets are dense by
moderately undercutting the bank, which provides hiding and resting
cover for fish .
VALUE FOR REHABILITATION OF DISTURBED SITES :
Richardson willow is useful in stabilizing streambanks and providing
erosion control on severely disturbed sites . Willow species are
the most important colonizers of disturbed sites in the Alaskan taiga
because of their ability to produce root and root crown shoots, which
provide for quick recovery [13,37]. At an Alaskan arctic coastal plain
site, Richardson willow colonized bare areas of tundra after removal of
debris that had been there for 30 years. High percentage cover occurred
on sites with favorable moisture and nutrient regimes . Richardson
willow was found to be a poor colonizer in areas where crude oil was
spilled; plant recovery and establishment were extremely slow on these
Willow planting, using stem cuttings, has been recognized as a valuable
tool for restoring riparian habitat. Restoration of riparian habitat
benefits a large number of wildlife species .
OTHER USES AND VALUES :
Richardson willow is an important nectar producer for bees . Tough,
flexible shoots of Richardson willow can be woven into baskets and
furniture . Native Americans used the broth from boiled bark for
sore throats and tuberculosis [2 ].
OTHER MANAGEMENT CONSIDERATIONS :
Richardson willow is an important source of browse for moose in Alaska.
If the management objective is to provide moose habitat and if
environmental manipulation of species composition is possible, then only
the growth of preferred species, such as Richardson willow, should be
BOTANICAL AND ECOLOGICAL CHARACTERISTICS
SPECIES: Salix richardsonii
GENERAL BOTANICAL CHARACTERISTICS :
Richardson willow is an erect, much-branched shrub usually forming dense
clumps 3 to 6 feet (1-2 m) tall, sometimes to 15 feet (4.5 m) [5,35].
Young twigs are stout and densely hairy; older twigs are glabrous. The
bark of Richardson willow is smooth . Shrubs are composed of light
wood that becomes brittle with age; a single trunk rarely survives 60
years . In silty loam containing much organic matter, the roots of
Richardson willow are numerous in the top 7.5 inches (19 cm) of soil,
but become less abundant at 9 inches (23 cm). In frozen ground, roots
of Richardson willow do not exceed 9 inches (23 cm), but as the ground
thaws roots will grow up to 17 inches (43 cm) deep .
RAUNKIAER LIFE FORM :
REGENERATION PROCESSES :
Sexual reproduction: Richardson willow is dioecious. The fruit of
these plants is contained in a capsule that splits in half to release
many seeds that are then dispersed by wind or water [11,33]. Optimum
seed production occurs between 2 and 10 years . Bees are the chief
pollinating agents . The seeds of Richardson willow are
short-lived, germinating immediately on moist surfaces . Seed
germination occurs over a broad temperature range, 41 to 77 degrees
Fahrenheit (5-25 degrees C). This appears to be a compensatory
mechanism due to the short seed life . Germination of Richardson
willow seeds occurs best in moist, exposed mineral substrates that
receive direct sunlight .
Vegetative reproduction: Richardson willow will sprout from the root
crown or basal stem . It will root readily from stem cuttings or
from root and stem fragments buried in moist soil. Damaged and cut
stems produce prolific sprouts from the stembase or root collar .
SITE CHARACTERISTICS :
In Alaska and northern Canada, Richardson willow is found in wet areas
such as heaths, riverbeds, and streams; it is also found in the open
tundra, in pingos, and in mountains to at least 5,578 feet (1,700 m)
[14,39]. In interior Alaska, Richardson willow occurs in glacial drift,
outwash deposition areas, and on old river floodplains with considerable
variation in habitat conditions .
Soils: Richardson willow grows best in moist, alluvial bottomlands but
is also found in well-drained sandy or gravelly substrates. The general
pH range for willows is 5.5 to 7.5 . Growth of Richardson willow is
severely reduced when water levels are maintained at or above the root
crown for extended periods .
Plant associates: Richardson willow is commonly associated with the
following species: quaking aspen, white spruce (Picea glauca), black
spruce (P. mariana), Alaska paper birch (Betula resinifera), feltleaf
willow (Salix alaxensis), diamondleaf willow (S. pulchra), netleaf
willow (S. reticulata), American green alder (Alnus crispa), Sitka alder
(A. fruticosa), bog birch (Betula glandulosa), lichens (Ericaceae spp.),
huckleberry (Vaccinium spp.), bluejoint grass (Calamagrostis
canadensis), bluegrass (Poa spp.), sedges (Carex spp.), and mosses
(Polytrichum spp.) [12,27].
SUCCESSIONAL STATUS :
Richardson willow is an early successional species on moist sites and,
once established, may persist in areas with frequent disturbances such
as fires or flooding [7,8]. It also becomes important in the later
stages of riparian succession. Successional studies have shown that
once silt accumulates, Richardson willow will become established quickly
. Richardson willow was the first shrub to invade flood meadows,
after grasses and horsetail, on sandy alluvium in the tundra .
Richardson willow has low shade tolerance and therefore loses dominance
on sites that are heavily forested or succeeded by more shade-tolerant
SEASONAL DEVELOPMENT :
Richardson willow flowers from May through July or August [6,20]. The
fruit ripens soon after flowering, followed by seed dispersal in early
to midsummer .
SPECIES: Salix richardsonii
FIRE ECOLOGY OR ADAPTATIONS :
Richardson willow sprouts rapidly from basal stems and roots after fire
[28,32]. It produces minute, hairy seeds that are easily disseminated
by wind, and which are important in reestablishment [21,32].
POSTFIRE REGENERATION STRATEGY :
Ground residual colonizer (on-site, initial community)
Secondary colonizer - off-site seed
SPECIES: Salix richardsonii
IMMEDIATE FIRE EFFECT ON PLANT :
Richardson willow is a fire-tolerant species that sprouts readily from
the root or root crown after being top-killed by fire . If soil
organic layers are completely removed by fire, then the roots of
Richardson willow will not be able to sprout .
DISCUSSION AND QUALIFICATION OF FIRE EFFECT :
PLANT RESPONSE TO FIRE :
Richardson willow is an early successional species on burned sites
because of its ability to sprout vigorously from the root crown or roots
following fire [20,36]. Invasion by willows after fire depends on the
time of year of the fire, weather, and the absence or presence of a
mineral seedbed . Richardson willow seeds need a nutrient-rich
mineral seedbed to germinate. The chance of Richardson willow
establishing after a fire lessens as available mineral soil seedbeds
become occupied by faster growing herbaceous species and mosses .
Fire severity can affect willow postfire recovery. High-severity fires
can damage the roots to the point of no recovery [20,38]. Following
low-severity fires most willows will recover quickly because of their
ability to send up new roots from the root crown . Intense burning
can completely kill willows .
DISCUSSION AND QUALIFICATION OF PLANT RESPONSE :
FIRE MANAGEMENT CONSIDERATIONS :
Prescribed fire is widely used as a wildlife management tool to
rejuvenate decadent willow stands and stimulate sprouting [11,25].
Early seral stage communities created by fire can increase the carrying
capacity of winter range for moose in interior Alaska . Recurring
fires within some parts of the boreal forest have allowed aspen and
willow to replace coniferous forests . The tendency of willows to
expand quickly following fires and other disturbances and to form dense
thickets inhibits natural regeneration of conifers . Prescribed
burning can reduce initial competition from willow in areas to be
planted with cultivated species .
SPECIES: Salix richardsonii
1. Argus, George W. 1973. The genus Salix in Alaska and the Yukon.
Publications in Botany, No. 2. Ottawa, ON: National Museums of Canada,
National Museum of Natural Sciences. 279 p. 
2. Arno, Stephen F.; Hammerly, Ramona P. 1977. Northwest trees. Seattle,
WA: The Mountaineers. 222 p. 
3. Bliss, L. C.; Cantlon, J. E. 1957. Succession on river alluvium in
northern Alaska. American Midland Naturalist. 58(2): 452-469. 
4. Bliss, L. C. 1988. Arctic tundra and polar desert biome. In: Barbour,
Michael G.; Billings, William Dwight, eds. North American terrestrial
vegetation. Cambridge; New York: Cambridge University Press: 1-32.
5. Brayshaw, T. Christopher. 1976. Catkin bearing plants of British
Columbia. Occas. Pap. No. 18. Victoria, BC: The British Columbia
Provincial Museum. 176 p. 
6. Densmore, R. V.; Neiland, B. J.; Zasada, J. C.; Masters, M. A. 1987.
Planting willow for moose habitat restoration on the North Slope of
Alaska, U.S.A. Arctic and Alpine Research. 19(4): 537-543. 
7. Dorn, Robert D. 1976. A synopsis of American Salix. Canadian Journal of
Botany. 54: 2769-2789. 
8. Ebersole, James J. 1987. Short-term vegetation recovery at an Alaskan
arctic coastal plain site. Arctic and Alpine Research. 19(4): 442-450.
9. Eyre, F. H., ed. 1980. Forest cover types of the United States and
Canada. Washington, DC: Society of American Foresters. 148 p. 
10. Garrison, George A.; Bjugstad, Ardell J.; Duncan, Don A.; [and others].
1977. Vegetation and environmental features of forest and range
ecosystems. Agric. Handb. 475. Washington, DC: U.S. Department of
Agriculture, Forest Service. 68 p. 
11. Haeussler, S.; Coates, D. 1986. Autecological characteristics of
selected species that compete with conifers in British Columbia: a
literature review. Land Management Report No. 33. Victoria, BC: Ministry
of Forests, Information Services Branch. 180 p. 
12. Hanson, Herbert C. 1953. Vegetation types in northwestern Alaska and
comparisons with communities in other arctic regions. Ecology. 34(1):
13. Henry, G. H. R.; Gunn, A. 1991. Recovery of tundra vegetation after
overgrazing by caribou in arctic Canada. Arctic. 44(1): 38-42. 
14. Hulten, Eric. 1968. Flora of Alaska and neighboring territories.
Stanford, CA: Stanford University Press. 1008 p. 
15. Kartesz, John T.; Kartesz, Rosemarie. 1980. A synonymized checklist of
the vascular flora of the United States, Canada, and Greenland. Volume
II: The biota of North America. Chapel Hill, NC: The University of North
Carolina Press; in confederation with Anne H. Lindsey and C. Richie
Bell, North Carolina Botanical Garden. 500 p. 
16. Kershaw, G. Peter; Kershaw, Linda J. 1986. Ecological characteristics of
35-year-old crude-oil spills in tundra plant communities of the
Mackenzie Mountains, N.W.T. Canadian Journal of Botany. 64: 2935-2947.
17. Kuchler, A. W. 1964. Manual to accompany the map of potential vegetation
of the conterminous United States. Special Publication No. 36. New York:
American Geographical Society. 77 p. 
19. Little, Elbert L., Jr. 1979. Checklist of United States trees (native
and naturalized). Agric. Handb. 541. Washington, DC: U.S. Department of
Agriculture, Forest Service. 375 p. 
20. Lotan, James E.; Alexander, Martin E.; Arno, Stephen F.; [and others].
1981. Effects of fire on flora: A state-of-knowledge review. National
fire effects workshop; 1978 April 10-14; Denver, CO. Gen. Tech. Rep.
WO-16. Washington, DC: U.S. Department of Agriculture, Forest Service.
71 p. 
21. Lutz, H. J. 1956. Ecological effects of forest fires in the interior of
Alaska. Tech. Bull. No. 1133. Washington, DC: U.S. Department of
Agriculture, Forest Service. 121 p. 
22. Lyon, L. Jack; Stickney, Peter F. 1976. Early vegetal succession
following large northern Rocky Mountain wildfires. In: Proceedings, Tall
Timbers fire ecology conference and Intermountain Fire Research Council
fire and land management symposium; 1974 October 8-10; Missoula, MT. No.
14. Tallahassee, FL: Tall Timbers Research Station: 355-373. 
23. Machida, Steven. 1979. Differential use of willow species by moose in
Alaska. Fairbanks, AK: University of Alaska. 97 p. Thesis. 
24. McCluskey, D. Cal; Brown, Jack; Bornholdt, Dave; [and others]. 1983.
Willow planting for riparian habitat improvement. Tech. Note 363.
Denver, CO: U.S. Department of the Interior, Bureau of Land Management.
21 p. 
25. Meidinger, D.; Lewis, T.; Kowall, R. 1986. Biogeoclimatic zones and
subzones of the northern portion of the Mackenzie Timber Supply Area,
British Columbia. In: Northern Fire Ecology Project: Northern Mackenzie
Timber Supply Area. Victoria, BC: Province of British Columbia, Ministry
of Forests. 44 p. 
26. Milke, Gary Clayton. 1969. Some moose-willow relationships in the
interior of Alaska. College, AK: University of Alaska. 79 p. Thesis.
27. Miquelle, Dale G.; Van Ballenberghe, Victor. 1989. Impact of bark
stripping by moose on aspen-spruce communities. Journal of Wildlife
Management. 53(3): 577-586. 
28. Parminter, John. 1984. Fire-ecological relationships for the
biogeoclimatic zones of the northern portion of the Mackenzie Timber
Supply Area: summary report. In: Northern Fire Ecology Project: Northern
Mackenzie Timber Supply Area. Victoria, BC: Province of British
Columbia, Ministry of Forests. 59 p. 
29. Petersen, Stephen F. 1989. Beekeeping under northern lights. American
Bee Journal. 129(1): 33-35. 
30. Ferguson, Dennis E.; Boyd, Raymond J. 1988. Bracken fern inhibition of
conifer regeneration in northern Idaho. Ogden, UT: U.S. Department of
Agriculture, Forest Service, Intermountain Research Station. 11 p.
31. Risenhoover, Kenneth L. 1989. Composition and quality of moose winter
diets in interior Alaska. Journal of Wildlife Management. 53(3):
32. McCune, Bruce. 1982. Site, history and forest dynamics in the Bitterroot
canyons, Montana. Madison, WI: University of Wisconsin. 166 p. Thesis.
33. Schopmeyer, C. S., tech. coord. 1974. Seeds of woody plants in the
United States. Agriculture Handbook No. 450. Washington, DC: U.S.
Department of Agriculture, Forest Service. 883 p. 
34. U.S. Department of Agriculture, Soil Conservation Service. 1982.
National list of scientific plant names. Vol. 1. List of plant names.
SCS-TP-159. Washington, DC. 416 p. 
35. Viereck, Leslie A.; Little, Elbert L., Jr. 1972. Alaska trees and
shrubs. Agric. Handb. 410. Washington, DC: U.S. Department of
Agriculture, Forest Service. 265 p. 
36. Viereck, Leslie A. 1973. Wildfire in the taiga of Alaska. Quaternary
Research. 3: 465-495. 
37. Viereck, Leslie A. 1975. Forest ecology of the Alaska taiga. In:
Proceedings of the circumpolar conference on northern ecology; 1975
September 15-18; Ottawa, ON. Washington, DC: U.S. Department of
Agriculture, Forest Service: 1-22. 
38. Viereck, Leslie A.; Schandelmeier, Linda A. 1980. Effects of fire in
Alaska and adjacent Canada--a literature review. BLM-Alaska Tech. Rep.
6. Anchorage, AK: U.S. Department of the Interior, Bureau of Land
Management, Alaska State Office. 124 p. 
39. Walker, Marilyn D.; Walker, Donald A.; Everett, Kaye R.; Short, Susan K.
1991. Steppe vegetation on south-facing slopes of pingos, central arctic
coastal plain, Alaska, U.S.A. Arctic and Alpine Research. 23(2):
40. Wolff, Jerry O. 1978. Burning and browsing effects on willow growth in
interior Alaska. Journal of Wildlife Management. 42(1): 135-140. 
41. ITIS Database. 2004. Integrated taxonomic information system, [Online].
Available: http://www.itis.usda.gov/index.html. 
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