©1988 R. B. Shaw
AUTHORSHIP AND CITATION:
Gucker, Corey L. 2006. Cercocarpus montanus. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/plants/shrub/cermon/all.html [ ].
Revisions: Rim Fire information and burn photo added on 2 June 2015.
Cercocarpus argenteus Rydb.
= C. montanus var. argenteus (Rydb.) Martin 
C. betuloides Nutt. [43,50,107,112,134,321]
= C. m. var. glaber (Wats.) Martin 
C. b. var. betuloides Torrey & Gray 
= C. m. var. glaber (Wats.) Martin 
C. b. var. blancheae (Schneid.) Little 
= C. m. var. blancheae (Schneid.) Martin 
C. b. ssp. macrourus
= C. montanus Raf. [47,132]
C. b. var. macrourus (Rydb.) Jepson [107,112]
= C. m. var. macrourus (Rydb.) Jepson 
C. breviflorus Gray 
= C. m. var. paucidentatus (Wats.) Martin 
C. b. var. eximius (Schneid.) 
= C. m. var. paucidentatus (Wats.) Martin 
C. minutiflorus Abrams
= C. montanus var. minutiflorus (Abrams) Martin 
C. m. var. flabellifolius (Rydb.) Kearney, Peebles 
= C. m. var. montanus Raf. 
NRCS PLANT CODE :
true mountain mahogany
The scientific name of true mountain-mahogany is Cercocarpus montanus Raf. (Rosaceae) [30,47,64,115,132,133,313,317].
Many varieties are recognized. Throughout this review, varieties will be identified using their scientific names.
Cercocarpus montanus var. argenteus (Rydb.) Martin [86,178,223], silverleaf mountain-mahogany
Cercocarpus montanus var. blancheae (Schneider) Martin , island mountain-mahogany
Cercocarpus montanus var. glaber (Wats.) Martin [86,114,115,133,223], birchleaf mountain-mahogany
Cercocarpus montanus var. macrourus (Rydb.) Martin , Klamath mountain-mahogany
Cercocarpus montanus var. minutiflorus (Abrams) Martin , true mountain-mahogany
Cercocarpus montanus var. montanus [47,178], alderleaf mountain-mahogany
Cercocarpus montanus var. paucidentatus (Wats.) Martin [178,223], shaggy mountain-mahogany
Distribution of varieties:
Of the true mountain-mahogany varieties, C. m. var. montanus is the most widespread .
Cercocarpus montanus var. blancheae occurs on the Channel Islands and in the Western Traverse
Range in California and is considered rare [112,177]. Plants Database
provides a distributional map of true mountain-mahogany and its varieties. See Powell  for descriptions
of the distributions of C. m. var. paucidentatus, C. m. var. glaber, and
C. m. var. argenteus in the Trans-Pecos region of Texas. Kearney and Peebles  provide
information on the distributions of C. m. var. paucidentatus and C. m. var. montanus
FRES21 Ponderosa pine
FRES28 Western hardwoods
FRES32 Texas savanna
FRES33 Southwestern shrubsteppe
FRES34 Chaparral-mountain shrub
FRES36 Mountain grasslands
FRES38 Plains grasslands
STATES/PROVINCES: (key to state/province abbreviations)
Alligator juniper-oneseed juniper (Juniperus deppeana-J. monosperma)/true mountain-mahogany-desert ceanothus (Ceanothus greggii)
Colorado pinyon (Pinus edulis)/true mountain-mahogany type; Gambel oak (Quercus gambelii), sandpaper oak (Q. pungens), Alligator juniper, and oneseed juniper phases recognized
Mexican pinyon (P. cembroides)-true mountain-mahogany-evergreen sumac (Rhus virens var. choriophylla)
Colorado pinyon/true mountain-mahogany/spreading phlox (Phlox diffusa) in southeastern Great Basin 
true mountain-mahogany type associated with Mexican pinyon in Chiricahua National Monument 
true mountain-mahogany scrub in the Santa Catalina Mountains 
desert ceanothus-true mountain-mahogany-littleleaf sumac (R. microphylla) on Fort Bowie National Historic Site 
Colorado pinyon/true mountain-mahogany in northern Arizona  and in southeastern Arizona
gray oak (Q. grisea)/true mountain-mahogany in southeastern Arizona 
shrub live oak (Q. turbinella)-true mountain-mahogany 
Toumey oak (Q. toumeyi)/grassland in Chiricahua National Monument 
alluvial scrub vegetation in coastal southern California 
desert chaparral in San Bernardino Mountains 
scrub oak (Quercus spp.)/true mountain-mahogany chaparral type in the Liebre Mountains 
island chaparral in canyons and on slopes of interior southeastern Santa Catalina 
mixed mesic form chaparral on northern aspects in southern California 
true mountain-mahogany/Eastern Mojave buckwheat (Eriogonum fasciculatum) chaparral in the Liebre Mountains 
blue oak/true mountain-mahogany/hoary bowlesia-San Francisco woodland-star (Bowlesia incana-Lithophragma affine) in southern San Luis Obispo and northern Santa Barbara counties 
interior live oak-canyon live oak (Q. wislizenii-Q. chrysolepis) chaparral woodland (Horton, in 
island oak (Quercus spp.) woodland
cismontane California juniper (J. californica) woodland and scrub
southern interior cypress (Arizona cypress, Tecate cypress (Cupressus arizonica, C. forbesii)) forest
bigcone Douglas-fir (Pseudotsuga macrocarpa)-canyon live oak forest 
Utah serviceberry-Wyoming big sagebrush (Amelanchier utahensis-Artemisia tridentata ssp. wyomingensis)-true mountain-mahogany-antelope bitterbrush-mountain snowberry (Purshia tridentata-Symphoricarpos oreophilus)
Utah serviceberry -true mountain-mahogany 
true mountain-mahogany/bluebunch wheatgrass (Pseudoroegneria spicata) in northwestern Moffat County 
Gambel oak-true mountain-mahogany 
shrub live oak-true mountain-mahogany 
Rocky Mountain juniper (J. scopulorum)/true mountain-mahogany in Roosevelt National Forest 
Utah juniper (J. osteosperma)/true mountain-mahogany-squaw apple (Peraphyllum ramosissimum) in White River National Forest 
Utah juniper-Colorado pinyon/true mountain-mahogany/bluebunch wheatgrass in northwestern Moffat County 
oneseed juniper/true mountain-mahogany-wax current (Ribes cereum)
Colorado pinyon/true mountain-mahogany 
Colorado pinyon-Utah juniper/true mountain-mahogany in White River National Forest 
ponderosa pine/true mountain-mahogany/big bluestem (Andropogon gerardii) 
true mountain-mahogany-antelope bitterbrush in Wasatch Mountains 
Rocky Mountain juniper-limber pine (P. flexilis) in south-central part of the state 
true mountain-mahogany/sideoats grama (Bouteloua curtipendula) 
Utah serviceberry-true mountain-mahogany shrublands; rare type 
ponderosa pine/mountain shrub and mountain shrub types near Yucca Mountain 
true mountain-mahogany-serviceberry-Mexican cliffrose-littleleaf mock orange (Amelanchier spp.-Purshia mexicana-Philadelphus microphyllus)-sparse shrub
true mountain-mahogany-littleleaf mock orange-fendlerbush (Fendlera rupicola)-skunkbush sumac-sparse shrub
true mountain-mahogany-shrub live oak-skunkbush sumac-sacahuista-Wright silktassel (Nolina microcarpa-Garrya wrightii)-sparse shrub
true mountain-mahogany-Pinchot juniper (J. pinchotii)-desert ceanothus-shrub live oak-Gambel oak × shrub live oak/mixed grass
true mountain-mahogany-gray oak-shrub live oak-skunkbush sumac-sparse shrub
true mountain-mahogany-shrub live oak-turpentine bush (Ericameria laricifolia)-Wright silktassel-Schott's yucca (Yucca schottii)-sparse shrub 
true mountain-mahogany/sideoats grama 
gray oak-true mountain-mahogany in southern New Mexico 
gray oak-true mountain-mahogany-banana yucca (Y. baccata)/mixed forb 
shrub live oak-true mountain-mahogany
oneseed juniper/true mountain-mahogany 
Mexican pinyon/Toumey oak-true mountain-mahogany-pointleaf manzanita (Arctostaphylos pungens)/mixed grass 
Colorado pinyon/true mountain-mahogany/nodding brome (Bromus anomalus) high elevation woodland in the Gila National Forest; Gambel oak phase recognized
Colorado juniper-alligator juniper/true mountain-mahogany-gray oak/mountain muhly (Muhlenbergia montana) in western portion of state; Parry's agave (Agave parryi) phase recognized 
oneseed juniper/true mountain-mahogany
Colorado pinyon/true mountain-mahogany 
Oregon white oak (Q. garryana)/true mountain-mahogany, a rare type in the interior valleys of southwestern Oregon 
western chaparral climax in the Black Hills 
true mountain-mahogany-fragrant sumac (Rhus aromatica) in Black Hills National Forest 
ponderosa pine-Rocky Mountain juniper/true mountain-mahogany-common snowberry/little bluestem (S. albus/Schizachyrium scoparium) in southwestern Black Hills 
true mountain-mahogany-desert ceanothus-Gambel oak × shrub live oak-sandpaper oak
smooth-leaf sotol (Dasylirion leiophyllum)-true mountain-mahogany in Guadalupe Mountains National Park (Warnock in )
sandpaper oak-true mountain-mahogany in Trans Pecos region 
true mountain-mahogany-antelope bitterbrush in Wasatch Mountains 
Colorado pinyon/true mountain-mahogany/Indian ricegrass (Achnatherum hymenoides) in the central Great Basin of southern Utah 
Colorado pinyon-Utah juniper/true mountain-mahogany in Manti-LaSal National Forest 
Douglas-fir (Pseudotsuga menziesii)/true mountain-mahogany in southern portions of Abajo Mountains and Aquarius Plateau 
true mountain-mahogany/Idaho fescue (Festuca idahoensis) 
true mountain-mahogany-antelope bitterbrush in Wyoming Basin 
true mountain-mahogany-fragrant sumac in Black Hills National Forest 
true mountain-mahogany/soapweed yucca(Y. glauca) in Cheyenne River Basin 
Utah juniper/true mountain-mahogany in southwest
Rocky Mountain juniper-ponderosa pine/true mountain-mahogany in Black Hills 
ponderosa pine/true mountain-mahogany 
Baja California Norte:
mixed chaparral vegetation north of Ensenada 
Photographer R. J. Shaw
GENERAL BOTANICAL CHARACTERISTICS:
This description provides characteristics that may be relevant to fire ecology, and is not meant for identification. Keys for identification are available (e.g. [30,47,64,112,114,115,133,178,301,313,317]).
Aboveground description: True mountain-mahogany is a shrub or less often a small tree 3 to 20 feet (1-7 m) tall [30,86,114]. Dayton  suggests that on "favorable" sites true mountain-mahogany can assume small tree stature. Few to many branches are produced, and the form is erect to spreading [112,178,200,262]. Stems measure 0.2 to 2 inches (0.5-5 cm) in diameter and are sturdy and stiff [151,301]. The main stem is typically less than 30 inches (80 cm) in diameter . Researchers noted lignotubers on some shrubs in Nuttall's shrub oak-dominated chaparral vegetation in southern California . True mountain-mahogany bark is thin and smooth [7,47,104]. True mountain-mahogany is considered long-lived , and stem diameter has been used to successfully age shrubs in Wyoming and Utah [34,37].
Deciduous, persistent, "mostly deciduous," and "occasionally evergreen" are all used to describe true mountain-mahogany [78,151,178,245,317]. Deciduousness is latitude and/or variety dependent. Little  indicates that C. m. var. var. paucidentatus and C. m. var. glaber are both evergreen, while Sampson and Jesperson  describe C. m. var. glaber as "occasionally evergreen." In the Sierra Nevada foothills of Tehama County, California, true mountain-mahogany loses 25% to 75% of its leaves each fall. Researchers think leaf loss may be a response to summer drought conditions and not freezing . Leaves of true mountain-mahogany live 11.4 to 12 months in southern California . In central Utah, true mountain-mahogany leaves have a 5- to 6-month lifespan .
Leaves are simple, alternate, and vary from thin to somewhat thick and firm [53,83,114,115,134,177]. Clusters of leaves are common on true mountain-mahogany's short spur-like branches [107,262]. Leaf shape is variable and ranges from lanceolate to obovate [114,115,200]. Often leaves are broadest at their midpoint . Leaf blades are small and commonly length is less that 2 times the width . Length ranges from 0.4 to 2 inches (1-5 cm), and width ranges from 0.2 to 1 inch (0.5-2.5 cm) [47,78,114,115,151,200]. Margins are entire or toothed [107,301]. Many describe margins that are toothed only on the upper 50% or 33% of the leaf [30,50,200]. Two to four teeth per centimeter of margin is described by Stephens . Leaves are only slightly, if at all, revolute .
True mountain-mahogany flowers are perfect and without petals [114,115,262]. Flowers occur singly or in small clusters of up to 12 on the upper portions of 2-year-old stems [50,112,151]. Clustered flowers are common in the axils of the short spur-like branches and 2 to 3 flowers together are most common [86,133,200,301]. Flower tubes measure 0.1 to 0.7 inch (3-17 mm) long [114,115,317]. The fruit is a hard, cylindrical achene coated with short hairs [83,86,114,115,151]. Achenes measure 0.3 to 0.5 inch (8-12 mm) long [47,104,262]. The average mass of true mountain-mahogany seed collected in southern California was 18 mg . As is characteristic for the genus, true mountain-mahogany achenes have a persistent, somewhat twisted, hairy style or tail [7,50,83]. Styles are 1 to 4 inches (3-10 cm) long [78,107,112,317].
Underground description: The true mountain-mahogany root system is described as "extensive" . Typically there are substantial tap and lateral roots . Vegetative regeneration occurs from the root crown and from rhizomes [47,146]. Likely root penetration and spread are related to soil and site characteristics. Baker and Korstian  describe 1 to several taproots extending almost straight down with minimal spread. Rootlets on the taproots persisted to "great" depths. Taproots reached over 8 feet (2.5 m) deep. Based on data collected in 5 separate studies, true mountain-mahogany roots averaged 44.5 inches (113 cm) and ranged from 15.7 to 59.8 inches (40-152 cm) deep in northern New Mexico. The large stem base gave rise to "stout lateral roots" .
In California's San Gabriel Mountains, root systems of 2 young true mountain-mahogany shrubs that averaged 5 feet (1.5 m) tall had few branches and spread widely. The site had shallow soils overlaying rock. The majority of roots were concentrated in the shallow soil layer. The longest root measured 12.3 feet (3.75 m); maximum penetration was 5 feet (1.5 m), and maximum radial spread was 9 feet (2.7 m) . From a 3-foot (1 m)-deep excavation in Nuttall's scrub oak-dominated chaparral in the Cleveland National Forest in California, researchers found that a substantial number of true mountain-mahogany roots grew vertically beyond 3 feet (1 m). Sixty-three percent of the roots were 16 to 39 inches (40-100 cm) deep. The site burned 9 years before and received its last herbicide application 5 years before the study. Researchers claimed that treated and untreated sites were not visually different .
True mountain-mahogany roots growing in 3 different soil types in central Colorado's Manitou Experimental Forest showed little difference in maximum penetration and lateral spread. In Chubbs stony loam soils described as calcareous, loose, and fertile, a 4-foot (1 m) tall true mountain-mahogany shrub had roots penetrating to a maximum depth of 5 feet (1.5 m) and spreading laterally to a maximum of 10 feet (3 m). In Edole soils that had weak profile development, low fertility, and hard granite at 4.5- to 6-foot (1.4-1.8 m) depths, roots of a 4.5-foot (1.4 m) tall true mountain-mahogany penetrated to a maximum depth of 5 feet (1.5 m) and spread a maximum of 8 feet (2.4 m). In skeletal soils described as "little more than disintegrated parent rock," a 5-foot (1.5 m)-tall shrub had maximum root penetration of 3.5 feet (1.1 m) and maximum root spread of 5 feet (1.5 m). Based on provided root system sketches, most lateral roots were concentrated in the top 1 to 1.5 feet (0.3-0.5 m) of soil. Edole soil sites were dominated by ponderosa pine and Douglas-fir with some scattered quaking aspen (Populus tremuloides). Chubbs soil sites were ponderosa pine dominated, and Skeletal soil sites were Douglas-fir dominated .
True mountain-mahogany plants can support nitrogen-fixing bacteria in root nodules. Young plants (most under 12 years) excavated from sites in southeastern and central Wyoming had nodules deep below the soil surface (>10 inches (30 cm)). Nodulated plants were limited to areas where soils were low in nitrate (< 1ug/g) and organic matter (<1%) . From 2 plants in northern New Mexico, Aldon  identified 3 endomycorrhizal species and determined that 53% of the roots supported endomycorrhizae.
Variability: All the true mountain-mahogany varieties are described and a key for distinguishing varieties is available in . The following references provide descriptions of 2 or more varieties.
|C. m. var. argenteus||[223,301]|
|C. m. var. blancheae|||
|C. m. var. glaber||[112,172,223]|
|C. m. var. macrourus|||
|C. m. var. montanus||[128,178]|
|C. m. var. paucidentatus||[128,172,178,223,301]|
True mountain-mahogany may hybridize with curlleaf mountain-mahogany and littleleaf mountain-mahogany when distributions overlap. True mountain-mahogany × curlleaf mountain-mahogany hybrids result only when true mountain-mahogany provides the ovule. Curlleaf mountain-mahogany typically flowers 2 weeks before true mountain-mahogany in overlapping habitats, and hybrids are only found in true mountain-mahogany stands. Rarely do true mountain-mahogany × curlleaf mountain-mahogany hybrids produce offspring [274,306]. True mountain-mahogany and littleleaf mountain-mahogany hybrids may be reciprocal in southeastern Utah. True mountain-mahogany × littleleaf mountain-mahogany hybrids are found in areas dominated by littleleaf mountain-mahogany .
True mountain-mahogany × curlleaf mountain-mahogany hybrids grow in a tree-like form and have persistent
leaves characteristic of curlleaf mountain-mahogany that are larger and have teeth like those characteristic
of true mountain-mahogany in Utah's Uinta Basin [83,274,317]. Seed of true mountain-mahogany × curlleaf
mountain-mahogany hybrids can be produced on the current year's growth, whereas true mountain-mahogany seed
is produced exclusively on 2-year-old branches . True mountain-mahogany × curlleaf mountain-mahogany
hybrids are susceptible to frost and drought, and often winter dieback is 50% or more . Rarely do true
mountain-mahogany × littleleaf mountain-mahogany hybrids produce flowers, and these hybrids are considered
RAUNKIAER  LIFE FORM:
True mountain-mahogany reproduces by sexual and asexual means. Establishment from seed requires favorable conditions and is periodic [222,236]. Vegetative sprouts are produced following damage [1,47,251,332] but are also produced in the absence of damage .
Pollination: Flowers are wind-pollinated [151,153].
Breeding system: True mountain-mahogany is monoecious [115,262].
Seed production: Seed production is variable in time and space and can be affected by browsing animals and climatic conditions. While Plummer and others  report that true mountain-mahogany produces abundant seed, Kitchen  indicates that a high percentage of seed produced is not viable. Seed crops according to Biswell and Gilman  are heavy some years and scant in others. True mountain-mahogany was considered a "prolific" seed producer in Madera County, California, where in an undisturbed mature stand 1,900,000 seedlings/acre emerged in a single season .
Resources can be partitioned by true mountain-mahogany. One branch may produce a large amount of seed, while an adjacent branch produces little or no seed but produces considerable vegetative growth .
Unbrowsed true mountain-mahogany shrubs produced many more flowers and seeds than browsed shrubs in the Wasatch and Uinta mountains of Utah. Flower and seed production for both browsed and unbrowsed shrubs were greatest when precipitation was greatest .
Seed dispersal: Seeds are dispersed by wind and small mammals [1,33,151]. Wind-dispersed seed can travel up to 450 feet (140 m) from the parent plant .
High-quality true mountain-mahogany seeds are typically those 1st detached from the plant . Seed shed earliest may be most vulnerable to predation. Likely the pattern of seed shed affects seedling germination and establishment.
Seed banking: A soft, thin seed coat and lack of seed dormancy suggest that true mountain-mahogany seed does not persist in the seed bank [1,27,28]. The absence of true mountain-mahogany seedlings on burned sites led researchers to suggest that seeds are not adapted to long-term survival in soil and/or are not resistant to fire . True mountain-mahogany was present in vegetation in the Huachuca Mountains of Cochise County, Arizona, but did not emerge from soils collected in same area. Abundance of true mountain-mahogany on the site was not reported. A small area of 3.04 inches² (19.6 cm²) of soil was sampled . On 10-year-old burned sites in southern California mixed chaparral, C. m. var. glaber occurred irregularly and had low live cover (<10%) but did not emerge from soil collected on the site .
Seed stored in an open warehouse in Utah was evaluated periodically for viability. Temperature extremes in the warehouse during the 25-year period ranged from a low of -21.8 °F (-29.9 °C) to a high of 101 °F (38.3 °C). Stored seed was collected from Utah's Ephraim Canyon. In the 1st five years of storage, germination ranged from 61% to 68%. Germination decreased to 46%, 25%, and 3% after 7 years, 10 years, and 15 years of storage, respectively. No seed germinated after 20 and 25 years of storage . Viability of true mountain-mahogany seed was significantly (p<0.05) better in cold than in hot or variable hot and cold environments. Germination of true mountain-mahogany seed after 6 years of storage at -5 to -10°F (-21 to -23 °C) was 84.4 %, at 36 to 44 °F (2.2-6.74 °C) was 81.6%, at 55 to 95 °F (13-35 °C) was 61.6%, and in a metal container kept outdoors at -15 to 105 °F (-26 to 41 °C) was 42% .
Germination: Requirements for successful seed germination differ depending on the locality of production. Adequate moisture is often correlated with successful germination. In chaparral vegetation, germination is best in mature stands . True mountain-mahogany seed produced in California germinates readily [1,27]. Seed collected in California germinated soon after planting in pure river sand kept in a greenhouse at 60 to 70 °F (16-21 °C) . However, some indicate that true mountain-mahogany seed requires 1 to 3 months of cold stratification . Germination of seed collected at more northerly locations in New Mexico increased with increased duration of stratification, and the improvement was greatest for seeds collected from the 2 highest elevation sites. Improvement was least for seed collected in the furthest south of the 6 northern sites. Stratification did not improve germination of seed collected from the southernmost latitudes .
Moisture, climate, and site conditions: Increased germination of seed collected in Wyoming, Colorado, and Utah was linked to moisture in several independent studies. Seeds collected in Wyoming's Laramie Basin had 80% germination after 48 hours of soaking in water. The germination percentage was lower (57%) after 96 hours at 41 °F (5 °C) but without soaking . True mountain-mahogany seed germination was poor with alternating moist and dry conditions. Seed was collected in native habitats that were not described. Germination was 10% under the moist-dry treatments, much lower than the maximum germination of 65% using other tests, which were not described .
Smith  found that site and climate affected true mountain-mahogany seed germination and seedling establishment in ponderosa pine/bunchgrass communities in Colorado's Pike National Forest. Greenhouse, nursery, and field studies were conducted. Seed was collected from sites with different soils. Precipitation between the beginning of April and the end of August affected seed viability. Achenes collected from northwestern slopes were heavier, had a greater percent filled, and had higher germination percentages than seed collected on southwestern slopes. The largest and most viable seeds were collected from 2 sites with the least fertile soils. Average germination was 33% in a dry year and 80% when precipitation was normal or above. Although shrubs on infertile sites produced the most viable seed, a higher percentage of seedlings survived in fertile than infertile soils. Survival was nearly equal on north and south aspects, but seedlings grew fastest and were most "vigorous" on southern slopes. Pocket gophers and cottontail rabbits were important seedling predators .
Germination of true mountain-mahogany seed collected from 9 populations in Utah and Colorado correlated with winter precipitation but not elevation. Germination was 15% without any pretreatment. Germination increased with moist, cold treatments. After 2 weeks of moist chilling, germination averaged 28.6%; after 4, 6, 8, and 12 weeks germination was 57.0%, 83.4%, 89.7%, and 98%, respectively. Researchers suggested that snowpack levels of the sites may have affected the length of moist chilling required for seeds .
Seed size: Large-sized seeds germinated best out of 5 seed lots collected in northern New Mexico. Seeds less than 0.3 inch (0.8 cm) long were considered small, seeds 0.4 inch (0.9-1 cm) long were classified as medium-sized, and large seeds were greater than 0.4 inch (1 cm) long. Seeds were stratified at 38 to 42 °F (3.3-5.6 °C) for 28 days, then kept moist at room temperature. Germination of small-, medium-, and large-sized seeds averaged 37%, 54%, and 69%, respectively. Germination differences were significant (p≤0.05). Seed size alone did not explain germination differences. The seed lot with the largest number of large-sized seeds had the lowest average germination percentage .
Heat treatments: True mountain-mahogany seed collected below 6,600 feet (2,000 m) in southern California germinated readily on moist filter paper after 1 month of stratification but was significantly (P>0.001) reduced by 5 minute heat treatments of 250 °F (120 °C). Light and dark treatments produced very similar germination percentages, and seeds moistened with a solution of charred chamise (Adenostoma fasciculatum) were not adversely affected .
Seedling establishment/growth: Successful true mountain-mahogany seedling establishment is often correlated with adequate moisture. In chaparral communities, seedling establishment is sporadic and likely restricted to high moisture years . Seedlings are vulnerable to frost and drought, and on harsh sites, establishment occurs only during "favorable" conditions . Keeley  indicates that seedlings require mesic conditions and a well-developed litter layer. During a wet winter in San Diego County, true mountain-mahogany seedlings were abundant although they were absent the previous 5 years . The number of seedlings and saplings varied from 0 to 36,530/ha in California chaparral vegetation that was unburned for 56 to 118 years. For more on this study, see Succession in chaparral . Brooks  found no true mountain-mahogany seedlings in vegetation types adjacent to true mountain-mahogany stands in Wyoming's Laramie Basin and suggested that soil differences between the distinct vegetation types may have affected seedling establishment. In undisturbed true mountain-mahogany stands in Madera County, California, 1,900,000 seedlings/acre were produced in a single season. Mortality, however, was 98% after 2 years .
Moisture and site conditions affect establishment and growth: Woodmansee  studied seedling establishment in northwestern and north-central New Mexico. Germination occurred in 1967 when summer rainfall was above normal but not in 1968 when summer rainfall was again above normal. Timing of the rainfall may have been a factor. Spring emergent seedlings did not survive. Seeds germinated on or just below the soil or litter surface. Just a small number of seedlings were observed from 1967 to 1969, but survival was high (50%-100%) for the 1st year. Surviving seedlings grew in areas that were shaded for most of the day by associated vegetation and where herbaceous vegetation was scarce.
In northwestern Nebraska, aspect affected seedling establishment. Seedlings established in populations on southwestern but not on northeastern slopes. When populations were visited 10 years later, the researcher indicated that plant growth was slow and that the number of stems per plant decreased with plant age . True mountain-mahogany seedling production, survival, and growth were monitored on an herbaceous-dominated site in the western foothills of Tehama County, California. Grasses were predominantly annuals and included soft chess (Bromus hordeaceus), red brome (B. rubens), and foxtail fescue (Vulpia myuros). Twenty-one C. m. var. glaber seedlings emerged in the spring of 1948; of those, 3 survived to the spring of 1960, and seedling height averaged 7.2 inches (18 cm) . Additional information from Smith  on seedling survival as it related to site conditions is available in Germination.
Postfire seedling establishment: Many report that true mountain-mahogany seedlings are rare in the early postfire years [139,146] and that seedling establishment is most typical in mature stands . Keeley  indicates that although seed production soon following fire can be substantial, seedlings are uncommon in chaparral communities less than 50 years old. For more information on postfire seedling production, see Seed survival and postfire seedling production.
Mature plant growth: Annual twig growth of browsed and unbrowsed shrubs was greatest when precipitation levels were greatest in Utah's Unita and Wasatch mountains .
Vegetative regeneration: True mountain-mahogany regenerates through asexual means following aboveground stem removal from browsing and/or fire; however, asexual sprouting also occurs in the absence of aboveground damage. Researchers noted lignotubers on some C. m. var. glaber plants in Nuttall's scrub oak-dominated chaparral vegetation in southern California . Root crown sprouting following fire is considered "vigorous" and "rapid" [1,33]. Sprouts from rhizomes (~3 feet (1 m) from the main stem) occur as well . Fire severity may affect postfire sprouting. Researchers found more and taller spouts in areas burned at low severity than on unburned sites or sites burned in high-severity fires in Colorado. Root crown and epicormic sprouts occurred on the low-severity burned sites. Only root crown sprouts occurred on high-severity burned sites, and only epicormic sprouts occurred on unburned sites .
Plants that received over 300% utilization, meaning current annual growth and previous year's growth were removed, compensated but did not overcompensate for removal. Browsing occurred for 4 years, but utilization of 300% was typical for only 1 year. True mountain-mahogany cover was often lower on browsed than protected sites, but the height of browsed and unbrowsed shrubs was similar. Animals using unprotected sites were cattle, moose, elk, antelope, and deer [288,289]. In southwestern Colorado, the current annual growth and old wood to 0.2 inch (4 mm) above ground were removed from true mountain-mahogany shrubs. Early in the study, shrubs produced new sprouts, but no plants survived more than 6 years of the severe clipping treatments .
Sites in the Silverado Canyon in the Santa Ana Mountains of California, which were unburned for at least 60
years, revealed stem age structures that suggested new stems sprouted in the absence of fire . True
mountain-mahogany produced sprouts from root crowns and from rhizomes ~3 feet (1 m) or more away from the
main trunk in California chaparral that was unburned for 56 to 120 years. The largest number of rhizome
sprouts occurred on sites overtopped by other vegetation . For additional information on sexual and
asexual recruitment in this area see Keeley  in Succession in chaparral.
True mountain-mahogany occurs on dry slopes, hills, ridges, mesas, desert foothills, and rocky outcrops. It is associated with chaparral, northern oak woodlands, pine (Pinus spp.)-oak woodlands, mixed coniferous forests, alpine shrub, mountain brush, sagebrush (Artemisia spp.), aspen, and pinyon-juniper (Pinus-Juniperus spp.) vegetation [7,47,104,112,114,200,234,262,301,317,321].
In southeastern Arizona, Mouat  found that true mountain-mahogany was restricted to high-elevation sites with hilly or mountainous terrain and a high density of drainages (mostly >6 mi/mi²). Slopes occupied were steep (averaging >45%), and north aspects were preferred . In the Wasatch mountains of north-central Utah, true mountain-mahogany stands occurred on slopes ranging from 25% to 60%. True mountain-mahogany stands on south-facing slopes were consistently found at higher elevations than stands on north-facing slopes, suggesting that a minimum moisture level requirement may exist in this area . True mountain-mahogany was significantly (p<0.01) associated with north aspects in the San Gabriel and San Bernardino mountains of southern California .
Climate: True mountain-mahogany's large distribution suggests a wide tolerance of climatic conditions but limits an exhaustive review of climatic tolerance. Kitchen  indicates that the annual precipitation in true mountain-mahogany habitats ranges from 9.8 to 26 inches (250-660 mm). However, the Oregon white oak/true mountain-mahogany type in interior southwestern Oregon receives only 1.6 inches (40 mm) of precipitation per year . Climate regimes for several true mountain-mahogany populations are provided.
California: San Mateo, coastal California, experiences a Mediterranean climate. The mean July and January temperatures are 68.2 °F (20.1 °C) and 48 °F (9.1 °C), respectively. Freezing occurs only occasionally. The average annual precipitation is 25.7 inches (654 mm), and a severe dry season of approximately 5 months (May to September) receives less than 0.59 inch (15 mm) of rainfall per month .
Colorado: Forty years of data collection in Mesa Verde, Colorado, revealed average annual precipitation of 18.3 inches (465 mm). Normal annual snow levels average 78 inches (2,000 mm), and May and June are dry months. In the late summer and early fall, brief intense storms are typical. The average monthly maximum temperature for a 40-year period was 102 °F (39 °C), and the minimum was -20 °F (-29 °C) .
New Mexico: In Carlsbad Caverns National Park in southern New Mexico, the climate is semiarid continental. Seventy-eight percent of annual precipitation occurs from May to October, and the average annual precipitation is 14 inches (360 mm). An extreme low of annual precipitation was 4.3 inches (110 mm) in 1951, and an extreme high of 43 inches (1,100 mm) was recorded in 1941. Droughts are common in southern New Mexico .
Utah: In true mountain-mahogany habitats of southeastern Utah, the climate is semiarid. Humidity levels are low, and precipitation is variable. The range in diurnal and annual temperatures is great; the average January and July temperatures are 27 °F (-3 °C) and 72.3 °F (22.4 °C), respectively. Annual precipitation averages 12.8 inches (324 mm) .
Wyoming: In Wyoming's Laramie Basin, the frequency of wind speeds greater than 2 miles (3 km)/hour in true mountain-mahogany stands are 77% to 87%, and wind speeds average 2.7 miles (4.3 km)/hour and 5.5 miles (8.9 km)/hour for 2 sites .
Elevation: True mountain-mahogany typically occupies habitats between 4,000 and 10,000 feet (1,200-3,000 m) throughout its range . Narrower elevational ranges by state and variety are provided below:
|State/region||Variety, if applicable||Elevation in feet|
|C. m. var. glaber||3,000-6,500|
|C. m. var. paucidentatus||5,000-8,000 |
|California||under 8,200 [112,200]|
|C. m. var. var. blancheae||below 2,000|
|C. m. var. glaber*||under 8,200|
|C. m. var. macrourus||2,600-5,900 |
|New Mexico||4,500-9,500 |
|C. m. var. argenteus||6,000-8,000|
|C. m. var. montanus||6,000-8,000|
|C. m. var. paucidentatus||6,000-8,500 |
|Texas (Trans-Pecos region)||C. m. var. argenteus||7,500-8,500|
|C. m. var. glaber||3,400-7,300|
|C. m. var. paucidentatus||4,600-8,000 |
|Utah (Unita Basin)||6,000-8,500 |
|Intermountain West||3,900-9,800 |
|C. m. var. argenteus (NM and TX)||4,000-8,500|
|C. m. var. paucidentatus (AZ, NM, TX, and Mexico)||4,500-8,500 |
Soils: True mountain-mahogany occupies sites with well-drained, coarse, poorly developed, very shallow to moderately deep soils with almost neutral pH [34,151,260,271]. In the north-central Plains, true mountain-mahogany occurs on eroded hillsides with "badland" soils . Habitats in north-central Utah have rocky, shallow soils with poor development . In southeastern Arizona, true mountain-mahogany occurs predominantly on nonalluvial soils where limestone is the primary parent material. Soils are often very shallow and rocky [198,311]. True mountain-mahogany was also absent from alluvial soils in the Laramie Basin of Wyoming. True mountain-mahogany is rare or absent on serpentine soils in California's Santa Ana Mountains . In Idaho true mountain-mahogany is found on poor shallow soils on rocky ridgelines but also on moister and deeper soils in the canyon bottoms . Based on studies in the Intermountain West, Brotherson  reports that true mountain-mahogany habitats have an average soil depth of 9.7 inches (24.5 cm), pH of 7.3, soluble salt levels of 426 ppm, and clay, silt, and sand percentages of 33.5, 37.7, and 28.7, respectively.
In the Wasatch Mountain range in north-central Utah, stands where true mountain-mahogany was a dominant or
codominant occurred on sites with soil depths of 3 to 19 inches (8-48 cm). Soils were classified as clay to
sandy clay loams, and pH was neutral to slightly alkaline. Density of true mountain-mahogany was positively
correlated with the percent clay and silt in the soil and negatively correlated with the percent sand .
In another study involving 20 north-central Utah sites, percent clay was positively correlated with true
mountain-mahogany height, density, frequency, and cover. This finding together with observations of true
mountain-mahogany at higher elevation sites on south slopes than on north slopes suggested that water retaining
properties were important in dictating true mountain-mahogany's distribution . In Utah's Uinta Basin,
soil depth beneath true mountain-mahogany plants averaged 7.5 inches (19 cm), and pH averaged 7.7. Soils
were 75% sand, 11% clay, and 14% silt .
True mountain-mahogany grows best in full sun , tolerates early-seral site conditions, and is dominant in several late-seral or climax communities . For additional information on late-seral or climax communities where true mountain-mahogany is dominant, see Habitat types and plant communities. Following disturbances in areas where true mountain-mahogany is established, colonization by this species is rapid.
In southern California's San Sevaine area, true mountain-mahogany occurs in both pioneer and mature alluvial scrub vegetation. Pioneer communities, in inactive stream channels or recently scoured stream beds, are characterized by sparse vegetation and low species diversity. Mature communities are dominated by shrubs and subshrubs . True mountain-mahogany is a common understory species in old-growth ponderosa pine forests in southwestern Colorado's San Juan National Forest. Old-growth stands are those that have passed through the stand initiation stage and stem exclusion stage of development . In Utah's Wasatch chaparral, true mountain-mahogany vegetation occurs on rocky ridges and represents a stage in xeric site succession and not a late-seral or climax vegetation type .
Succession in chaparral: Succession following disturbances in chaparral vegetation involves little or no compositional change . The typically self-replacing recovery in this vegetation type has been termed autosuccessional. Shrubs present in mature stands, including true mountain-mahogany, are normally present in the 1st postfire year .
Several studies report abundance of true mountain-mahogany in different aged chaparral communities. In chaparral communities in the San Gabriel and San Bernardino mountains, true mountain-mahogany cover was 6.5% in 2- to 8-year-old stands, 15.5% in 9- to 21-year-old stands, 4.6% in 22- to 40-year-old stands, and 4.2% in 41- to 96-year-old stands . Researchers suggested that true mountain-mahogany may be reduced by taller shrubs such as Nuttall's scrub oak, hollyleaf redberry (Rhamnus ilicifolia), and/or hollyleaf cherry (Prunus ilicifolia) when chaparral stands approach 50 years old . However, true mountain-mahogany density was greater in ancient (130-150 years unburned) than mature (50-60 years unburned) chaparral vegetation in the Sierra Nevada. Density in mature stands was 209/ha and in over-mature stands was 487/ha .
Succession in pinyon-juniper: True mountain-mahogany typically appears in the shrub/grass stage of postfire succession in pinyon-juniper communities. This stage can appear immediately following fire and cover may reach prefire levels within 7 years if pinyon-juniper cover was low (10% to 15%) when burned. If pinyon-juniper cover was 60% or greater when burned, sprouting shrub cover may be less than 2% in the 10th postfire year . In Daggett County, Utah, true mountain-mahogany coverage was compared in different-aged Colorado pinyon-Utah juniper stands. True mountain-mahogany cover decreased with increased pinyon-juniper canopy cover which generally increased with stand age. A summary of the results is presented below :
|Approximate stand age (years)||70||150||215||140|
|Colorado pinyon-Utah juniper cover (%)||18.5||29.2||53.9||62.1|
|True mountain-mahogany cover (%)||29.6||12.7||0.9||0|
Postfire succession: Many studies have compared true mountain-mahogany abundance on sites with different fire histories and increasing time since fire. True mountain-mahogany abundance may vary with time since fire, fire history, site conditions, and/or prefire stand characteristics.
True mountain-mahogany coverage varied on sites with different recent fire histories and substrates in California's Santa Monica Mountains. All sites burned 4 years before measurements were made. Substrates included andesite, granite, and sandstone. On sites that had burned 22 years earlier, the highest coverage was 82.2%. On sites that burned 75 years before the most recent fire, the highest coverage was 29%. The inclusion of different substrates and lack of prefire or unburned comparisons makes assessing the importance of fire regime difficult .
In cis-montane chaparral vegetation of San Diego County, California, true mountain-mahogany frequency and cover were relatively unchanged from 1930 to 2001. Coverage and frequency were 3% and 10%, respectively, in 1930. Between 1930 and 2001, 41% of measured plots burned once and 40% burned 2 or more times. In 2001, true mountain-mahogany coverage was 3% to 4% and frequency was 13% .
True mountain-mahogany was present on all sites in a chronosequence ranging from 4-year-old burned sites to a 400-year-old climax Colorado pinyon-Utah juniper/mountain brush woodland in Mesa Verde National Park, Colorado. True mountain-mahogany cover, 14%, and frequency, 28%, were greatest in ~400-year-old climax stands. True mountain-mahogany cover and frequency were 1% and 6% on 30-year-old burned sites and 2% and 10% on 90-year-old burned sites. Presence was indicated on 4-year-old burned sites, but values were not reported. Most sites burned in summer fires, but other fire characteristics were not reported [67,68].
Importance of true mountain-mahogany generally increased as time since fire increased in mixed conifer forests in the White Mountains of south-central New Mexico. Importance values were the sum of relative density, relative basal area, and absolute frequency. True mountain-mahogany was absent from sites burned 3 years earlier, but importance values were 50, 104, 75, and 104 on 16-, 21-, 27-, and 80-year-old burned sites, respectively. Researchers reported that shrub stages could persist for up to 80 years and that substantial increases in conifers took 50 to 100 years .
The successional recruitment of true mountain-mahogany in California chaparral was studied in 50- to 120-year-old stands. The percentage of dead basal cover was greatest (34.1%) at the oldest site. True mountain-mahogany basal coverage and density were high in both the 74- and 83-year-old stands. Seedling and sapling production were absent in the oldest and youngest stands. Seedling density was by far the greatest in the 83-year-old stand, which authors reported was open and disturbed. Time since fire was not a good predictor for seedling and/or sapling growth. Data collected during this study are summarized below [140,141].
|Site||Pacific View||Bartlett Spring||Sequoia||Silverado|
|Time since fire (years)||56||74||83||118|
|Latitude||34° 05'||39° 09'||36° 28'||33° 45'|
|Basal coverage (m²/ha)||3.24||8.47||13.92||1.32|
|Stems produced in 1st decade after fire (%)||<1||0||4||0|
|Stems produced in most recent decade (%)||92||67||85||61|
|State/region||Flowering dates||Fruiting dates|
|Baja California Norte||March-April ||----|
(Santa Monica Mountains)
|Nebraska||early June||August |
|New Mexico||May-August ||----|
|South Dakota||early June||August |
|Utah (Uinta Basin)||May-June ||----|
|Great Plains||March-June ||----|
|Intermountain West||mid-May-June ||----|
|Pacific Northwest coast||February-June ||----|
Flowering and fruit production dates for true mountain-mahogany varieties are summarized below :
|C. m. var. argenteus||June-July|
|C. m. var. blancheae||March-June|
|C. m. var. glaber||February-June; rarely September-October|
|C. m. var. macrourus||June|
|C. m. var. minutiflorus||March-May|
|C. m. var. montanus||May-July|
|C. m. var. paucidentatus||May-November|
The following table summarizes the phenology of true mountain-mahogany based on 9 years of data collected at 2 elevations in Utah's Ephraim Canyon. Presented are the average dates of appearance :
|7,480||April 30||May 28||May 28||July 23||August 11||October 14|
|7,890||May 3||June 5||June 7||August 2||August 3||October 19|
Fire regimes: The true mountain-mahogany fire regime is dictated by its associated vegetation and distribution, both of which vary widely. In singleleaf pinyon, California juniper, and western juniper (Juniperus occidentalis) woodlands of the San Bernardino Mountains where true mountain-mahogany is common the fire cycle is an estimated 480 years . True mountain-mahogany is also a common shrub in mixed conifer and pinyon-juniper forests of New Mexico's El Malpais National Park where low-severity surface fires were common from ~1700 to 1880. An evaluation of 9 sites revealed an average fire return interval of 5 to 12 years .
Numerous studies of past and present fire regimes have been conducted in true mountain-mahogany habitats. Presented below is information on the fire history, fuel availability, and ignition sources and/or ignition probability in pinyon-juniper woodlands, mountain shrublands, mixed conifer forests, ponderosa pine forests, oak woodlands, and chaparral communities.
California chaparral: The fire regimes of southwestern chaparral, especially in southern California, have been investigated extensively. The lack of fire scars to reconstruct the fire history of these shrublands requires the use of alternate study methods. Fuel studies, ignition probabilities, postfire recovery of the vegetation, and ethnocentric histories are typically used to reconstruct the fire regimes. Using these methods there are still competing schools of thought about fire in chaparral systems.
Mature chaparral vegetation in southern and central California burns in stand-replacing fires. Rarely are herbaceous fuels sufficient to support surface fires, except maybe in the case of recently burned shrublands . Ethnohistories indicate that the native people of California burned mixed chaparral vegetation primarily in the fall, but some spring burning occurred as well. "Spot burning" was routine and likely created and maintained openings and ecotones. A range of successional stages and a complex of habitats were probably normal under these burning practices .
California chaparral vegetation is comprised of a mixture of species, including those that sprout following fire but require a fire-free period for successful seedling establishment and those that require fire for seed scarification and successful seedling establishment. While a fire return interval of 20 years would be fine for species that germinate with fire, it is at odds with species requiring a fire-free period for the recruitment of new individuals. This conundrum suggests that the "long-term stability of chaparral communities may require a mosaic of fire frequencies" . Not only does the fire return interval affect species recruitment; it may also affect species recovery. When chaparral vegetation burns frequently (~25 year fire return interval), the number of dead shrubs is typically low and fire severity will likely be low. Under these conditions, the potential for postfire sprouting is high and there are only small openings available for seedlings. When chaparral vegetation experiences long fire-free intervals (~100 years), there are more dead shrubs at time of the fire, and fire severity increases. Fewer postfire sprouts and increased mortality create larger openings on which seedlings may capitalize .
Postfire root crown and rhizome sprouting allow true mountain-mahogany to tolerate fire return intervals of 10 to 100 years. However, a fire-free interval of 100 years or more may be required for the successful recruitment of new individuals . When considering all the species present in chaparral communities, Keeley and others  indicate that long fire-free periods are tolerated. Based on biological requirements and fire adaptations of chaparral species, Zedler  suggests that there is not an "urgent need to reduce the time between fires" in chaparral systems. See Fire management considerations for recommendations on managing fire in chaparral communities.
Competing philosophies of thought on California chaparral fire regimes: There are 2 prevailing philosophies about the best way to view and estimate natural fire regimes in California chaparral. One "deterministic" approach proposes that fire frequency is driven by fuel loadings and that ignitions are not limiting. Under this "school of thought," chaparral vegetation may burn every 15 to 20 years. The other philosophy suggests that fire regimes are dictated by a series of stochastic events, especially unpredictable ignitions. Proponents of this philosophy believe that "chaparral is resilient to a wide range of fire frequencies" .
Minnich, a proponent of the "deterministic" theory, suggests that fire regimes in southern California chaparral have changed with 20th century fire exclusion. While the fire return interval of approximately 40 to 60 years has not changed considerably with fire exclusion practices, fire size and behavior have changed . Minnich describes an "erratic smolder and run" fire behavior before the 20th century. Behaviors of 3 chaparral fires that burned in 1896, 1898, and 1900 in the San Gabriel Mountains were studied from written accounts. Fires burned for 2 to 3 months during summer droughts. However, fire size was small (5,800 to 15,800 acres (2,300-6,390 ha)), suggesting that fire spread was slow, and fireline intensity was low. The smolder and run behavior that created a mosaic burn pattern and a community of fragmented fuels was created through the flare up of smoldering logs, trees, or other large fuels and the movement of embers into areas beyond the fire zone . With the exclusion of fires and greater fuel uniformity, fires are now large and severe . Since 20th century fire exclusion, fires are brief (< a few days) and extensive (>25,000 acres (10,000 ha)) . Minnich and Bahre  found further support for these fire regime theories in a study comparing chaparral vegetation on the California-Baja California Norte.
Keeley, a supporter of the "stochastic" theory, suggests evidence that young southern California chaparral stands effectively suppress fire spread is lacking. Weather, according to stochastic theory supporters, is the primary controller of fire spread . Santa Ana winds of southern California can exceed 60 miles (100 km)/hour and may have humidity levels below measurable levels. These winds create extreme fire conditions [97,142] that are not mitigated by stand age and/or species composition. Early 1900 fire exclusion practices have not decreased the fire frequency in southern California shrublands. Fire frequency has increased with increased human population. The current fire return interval for southern California shrublands is 30 to 40 years . A study in San Diego County mapped 20th century fire data on top of corresponding vegetation types. The resulting picture supported Keeley's thoughts that fire size has decreased while amount of area burned has increased over time; it did not support Minnich's theory that the number of fires per decade has decreased while fire intensity and size have increased .
Regardless of what past fire regimes in chaparral may have been, human influences on fire regimes and disturbance patterns in heavily populated areas are now a dominant factor shaping southern California shrublands. In Pinnacles National Monument, the fire record from 1934 to 1978 indicates that the majority of fires were human caused, and in southern California in general, 75% of fires were human caused . Keeley and Keeley  report that approximately 5% of California chaparral ignitions are natural. The introduction of persistent annual Mediterranean grasses has increased the probability of frequent fires . For more on how humans, as ignition sources and fire suppressors, fit into the 2 prevailing theories on chaparral fire regimes, see .
Arizona chaparral: Researchers evaluated the fire history of a shrub live oak-true mountain-mahogany chaparral community from 7 fire-scarred stumps from a surrounding 215 (87 ha)-acre ponderosa pine forest in central Arizona's Bradshaw Mountains. There were no living trees with fire scars, and all living trees were less than 100 years old. Stumps recorded 92 fires in the 174-year period from 1700 to 1874, suggesting that ~1 fire burned every ~2 years. However, fire spread is poor in this chaparral type when stands are less than 10 years old and grass cover is scant. Flammability increases each year without fire. Researchers suggest that, although the surrounding ponderosa pine forest burned on average every 2 years, chaparral communities likely did not burn at an interval of less than 15 to 20 years. Fire protection and grazing likely both played a part in the dramatically decreased fire frequency since the late 1800s in the area .
Oak woodlands: The fire history of oak woodlands in the Yosemite Wilderness was determined for the 1930 to 1983 time period. Woodlands with an overstory of canyon live oak, interior live oak, and gray pine and true mountain-mahogany common in the understory contributed 4.2% to the wilderness' vegetation. Just 1.9% of Yosemite's fires occurred in this vegetation type. The average size of fires greater than 10 acres (4 ha) was 177.5 acres (71.8 ha). While weather conditions could likely support a fire in any year, lightning strikes were infrequent. The average number of fires in this vegetation type for the 53-year period was 22 fires per year per million acres. Fires were considered "intense" .
Parsons  summarized the recent fire history of foothill chaparral and oak woodland communities found below 6,000 feet (1,830 m) in Sequoia National Park. From 1920 to 1929, a total of 10,350 acres (4,189 ha) of the approximately 69,000 (28,000 ha)-acre study area burned. From 1930 to 1978, a total of 105 lightning fires burned 1,460 acres (592 ha), and 107 man-caused fires burned 6,946 acres (2,811 ha). The majority of fires (91%) were less than 7 acres (3 ha). Oak savannah and chaparral had the most burned area from 1920 to 1978 .
Ponderosa pine forests: The fire history of Kings Canyon National Park in California was reconstructed from 10 fire-scarred trees over a 400 (160 ha)-acre study area. True mountain-mahogany was an occasional species in the ponderosa pine- and Jeffrey pine (Pinus jeffreyi)-dominated park. From 1775 to 1909, a fire burned somewhere in the study area on average once every 3.5 years, and the average fire interval per individual tree was approximately 11 years. There were no fire scars recorded on any tree after 1909 . In Yosemite National Park, the number of lightning strikes from 1985 to 1990 in low- and mid-elevation sites, where chaparral and ponderosa pine are typical, was significantly (p<0.05) lower than what would be expected for low- and mid-elevation sites based on chance .
Mixed dry conifer forests: An extensive study of El Malpais National Park in New Mexico revealed that frequent low-severity surface fires were common in the presettlement period of ~1700 to 1880. True mountain-mahogany was a common shrub in the mixed Douglas-fir, ponderosa pine, and pinyon-juniper forests. Nine sites were studied, and on average, sites burned every 5 to 12 years. The minimum and maximum fire return intervals were 1 and 55 years. From before 1700 to 1782, fires were frequent, patchy, and occurred predominantly in the late season during drought years. From 1783 to 1794, there were almost no fires, which was likely due to increased rainfall during this period. From 1795 to 1880, fire intervals were longer than in the pre-1700 to 1782 time period. Fire size was larger, and early season fires were most typical. Fires were uncommon or absent again from 1881 to 1892, and while ignitions occurred from 1893 to 1939 fire spread was limited. Intense domestic sheep grazing, which reduced fine fuel availability, was likely partially responsible for fewer fires from the late 1800s to 1939. From 1940 to 1992, the fire-free intervals were the longest in the past 500 years. Grazing, fire exclusion, logging, and/or road building affected the likelihood of fire after 1940. Large fires were reported in 1976 and 1989. The researcher indicated that the lack of fire in this area has created fuel and vegetation conditions that will likely support uncharacteristic high-severity, stand-replacing fires .
Dry Douglas-fir/true mountain-mahogany habitats in Utah likely did not burn frequently due to the discontinuity of fine fuels. However, when fires did burn they were likely severe . Before fire exclusion practices, areas in the San Bernardino Mountains dominated by bigcone Douglas-fir were characterized by low-severity, summer-long fires. The interval between stand-replacing fires was likely long. Present stand replacement due to fires is ~26% and seemingly "excessive" when compared with the historic spatial extent of stand establishment .
Pinyon-juniper/petran chaparral: Using stand age structure sampling, researchers reconstructed the fire history for Colorado pinyon-Utah juniper-Rocky Mountain juniper, and petran chaparral for the 1850 to 2000 time period in Colorado's Mesa Verde National Park. Gambel oak, Utah serviceberry, and true mountain-mahogany are typical in petran chaparral. The sampling method allowed detection of only large severe fires. The fire cycle for the petran chaparral was an estimated 100 years. The fire cycle for the pinyon-juniper vegetation was ~400 years. Fire cycles for the early 1900s were long (~200 years for chaparral and ~1200 years for pinyon-juniper), likely due to livestock grazing, successful exclusion efforts, and/or a moist climate that prevailed in the southwest during this time. The fire regime since ~1950 more approximates that of the late 1800s. Changes in the fire regime may have been due to the removal of grazing and/or failed suppression efforts during severe weather conditions. From 1926 to 1977, the park averaged 8 fires per year; 90% of the fires were lightning ignited. Only 2% of the fires were greater than 10 acres (4 ha) but account for greater than 95% of the area burned in the 1900s. Large stand-replacing fires occurred during periods of prolonged drought and high wind conditions. Low-severity surface fires have been of limited extent in Mesa Verde Park .
Fire regimes in singleleaf pinyon, California juniper, western juniper forests of the San Bernardino Mountains have not changed considerably with the exclusion of fire. Fire return intervals were long before and after exclusion. Long fire return intervals are attributed to low productivity and fuel build up rates . Researchers estimate the fire cycle for these pinyon-juniper woodlands at 480 years. Fires were stand replacing and typically small. Postfire succession proceeds slowly in these woodlands .
Mountain shrublands: Bare ground and sparse grass and forb growth in most true mountain-mahogany stands in Wyoming's Laramie Basin  suggest that fires are uncommon.
The following table provides fire return intervals for plant communities and ecosystems where true mountain-mahogany is important. For further information, see the FEIS review of the dominant species listed below.
|Community or Ecosystem||Dominant Species||Fire Return Interval Range (years)|
|California chaparral||Adenostoma and/or Arctostaphylos spp.||<35 to <100|
|sagebrush steppe||Artemisia tridentata/Pseudoroegneria spicata||20-70 |
|Wyoming big sagebrush||Artemisia tridentata var. wyomingensis||10-70 (x=40) [300,328]|
|coastal sagebrush||Artemisia californica||<35 to <100 |
|plains grasslands||Bouteloua spp.||<35 [217,324]|
|California montane chaparral||Ceanothus and/or Arctostaphylos spp.||50-100 |
|curlleaf mountain-mahogany*||Cercocarpus ledifolius||13-1,000 [13,248]|
|mountain-mahogany-Gambel oak scrub||Cercocarpus ledifolius-Quercus gambelii||<35 to <100|
|Ashe juniper||Juniperus ashei||<35|
|western juniper||Juniperus occidentalis||20-70|
|Rocky Mountain juniper||Juniperus scopulorum||<35|
|pinyon-juniper||Pinus-Juniperus spp.||<35 |
|Rocky Mountain bristlecone pine||P. aristata||9-55 [62,63]|
|Mexican pinyon||Pinus cembroides||20-70 [192,276]|
|Rocky Mountain lodgepole pine*||Pinus contorta var. latifolia||25-340 [19,20,279]|
|Sierra lodgepole pine*||Pinus contorta var. murrayana||35-200 |
|Colorado pinyon||Pinus edulis||10-400+ [72,85,136,217]|
|interior ponderosa pine*||Pinus ponderosa var. scopulorum||2-30 [10,15,168]|
|Arizona pine||Pinus ponderosa var. arizonica||2-15 [15,45,249]|
|quaking aspen (west of the Great Plains)||Populus tremuloides||7-120 [10,93,183]|
|mountain grasslands||Pseudoroegneria spicata||3-40 (x=10) [9,10]|
|Rocky Mountain Douglas-fir*||Pseudotsuga menziesii var. glauca||25-100 [10,11,12]|
|coastal Douglas-fir*||Pseudotsuga menziesii var. menziesii||40-240 [10,197,238]|
|California oakwoods||Quercus spp.||<35 |
|oak-juniper woodland (Southwest)||Quercus-Juniperus spp.||<35 to <200 |
|canyon live oak||Quercus chrysolepis||<35 to 200|
|blue oak-foothills pine||Quercus douglasii-P. sabiniana||<35|
|Oregon white oak||Quercus garryana||<35 |
|California black oak||Quercus kelloggii||5-30 |
|interior live oak||Quercus wislizenii||<35 |
|little bluestem-grama prairie||Schizachyrium scoparium-Bouteloua spp.||<35 |
|True mountain-mahogany sprouts on the Stanislaus National Forest, 15 months after the 2013 Rim Fire. Photo by Becky Howard.|
Seed is typically killed by fire, and it is rare to find seedlings in early postfire communities . Biswell and Gilman  indicate that late season fires are most detrimental to seedling production on burned sites. Hanes and Jones  report that seedlings may establish from off-site seed sources in early postfire communities, although Halsey  indicates that true mountain-mahogany seed germination is best in mature stands of chaparral.
Postfire sprouting: Fire timing, fire severity, site conditions, and ecotypic variation may affect true mountain-mahogany postfire sprouting. Ferguson  suggests that the true mountain-mahogany postfire sprouting response may be ecotype dependent. Hanes  indicates that true mountain-mahogany sprouts are typically taller on burned desert than on burned coastal sites.
Production of postfire sprouts, however, is not guaranteed. On 1- to 2-year-old burned sites in the San Gabriel Mountains density of sprouting true mountain-mahogany shrubs was 2/acre and nonsprouting shrub density was 3/acre. Information about the fire was lacking. Fire severity, seasonality, or behavior may have affected sprouting .
Sprouting was vigorous after a fire in the western foothills of the Sierra Nevada Mountains of Tehama County. The fire did not kill true mountain-mahogany shrubs, and postfire sprouts produced seed in the 5th postfire year. Researchers noted that fire timing affects true mountain-mahogany's sprouting response. If burned in a late season fire (late September or after), postfire sprouts may not appear until the following spring. If burned during the growing season, postfire sprouts typically appear by postfire week 3. Heavy browsing of postfire spouts frequently kills true mountain-mahogany .
Sixty-four percent of true mountain-mahogany shrubs sprouted ~4.5 months after an "extremely hot" July fire in the San Gabriel Mountains. A greater percentage of plants were sprouting on the low-elevation site (2,100 feet (640 m)) than at the high-elevation sites (3,100 (940 m) and 4,100 feet (1,200 m)). Some areas had been sprayed with herbicide prior to the fire; whether or not this affected postfire sprouting was not discussed .
Researchers found that true mountain-mahogany produced more and taller spouts in areas burned at low severity than on unburned or high-severity burned sites following the Hi Meadow Fire in Colorado. Basal and epicormic sprouts occurred on low-severity burned sites. Only basal sprouts occurred on high-severity burned sites and only epicormic sprouts on unburned sites . Similar results were found following a January fire in Carlsbad Caverns National Park. More and longer sprouts were produced by "lightly" burned or scorched true mountain-mahogany shrubs than by shrubs that were "burned to char." Sprouts averaged 2 inches (5 cm) in postfire month 2 and 8 inches (20 cm) in postfire month 6. Shrubs regained their prefire height of 2.5 feet (0.76 m) three and a half years following the fire .
Seed survival and postfire seedling production: Heat treatments decrease true mountain-mahogany germination, but seeds have survived fire and seedlings have appeared in early postfire communities. However, the recruitment of new true mountain-mahogany shrubs may require a fire-free interval of 100 years or more , and seeds are thought to germinate best in mature chaparral stands . For information on heat treatment effects on true mountain-mahogany seed see Heat treatments.
Seeds in the canopy of tree-like true mountain-mahogany plants survived fire in the foothills of Madera County, California. Fourteen- to sixteen-foot-tall trees burned in a prescription fire in July of 1949. Some trees were heavy with seed at the time of the fire. Trees were killed but seeds were not. Three days after the fire, true mountain-mahogany seedlings were dense. In the summer of 1951, there was on average a 2-year-old seedling per every 3 ft² (0.3 m²). Researchers indicated that seed survival in the canopy would be unlikely during an "intense summer" fire .
Following fall fires in southern California, seedlings were present as soon as the 2nd postfire year on chaparral sites. A total of 200± 100 (sx) true mountain-mahogany seedlings occurred on burned chaparral sites. No seedlings occurred on burned sage-scrub sites. No seedlings occurred in the 1st or 3rd postfire years. Approximately 33% of the total number of seedlings occurred equally in the 2nd, 4th, and 5th postfire years. Seedling survival was not reported .
True mountain-mahogany cover was reduced, but seedling density increased following fire in a mixed chaparral/oak community with an annual herbaceous understory. Before the fire, interior live oak, which covered 60% of the plot, was selectively crushed and piled. This site preparation decreased true mountain-mahogany cover from 3% to 2%, and 1 and 2 years following the fire, cover was nearly zero. Seedlings were absent before the fire but 2 years after the fire there were 13 true mountain-mahogany seedlings in the plot. Fire season, conditions, or characteristics were not described .
Eighty-three percent of the postfire true mountain-mahogany population was sprouts on a low-elevation 1,800 feet (560 m) site in southern California. Postfire populations were 100% sprouts on mid- (3,300 feet (1,000 m)) and high- (5,480 feet (1,670 m)) elevation sites in the same area. Average postfire sprout heights were 27 inches (69 cm), 18 inches (45 cm), and 12 inches (31 cm) at the low-, mid-, and high-elevation sites, respectively. Seedling heights on the low-elevation site were not reported .
See Fire Studies in FEIS for further information on prescribed fire and postfire responses of many plant species. These Fire Studies are available for birchleaf mountain-mahogany:
Studies lacking information on fire behavior or burned and unburned comparisons: True mountain-mahogany sprouts required over 2 months recovery time after a July fire in southern California's Anza-Borrego Desert State Park. Sprout and shrub number and productivity of C. m. var. glaber were evaluated at postfire months 2, 4, 7, and 10. An estimated 25% of true mountain-mahogany shrubs suffered mortality. There were no sprouts in postfire month 2, and the density of sprouts and productivity of shrubs decreased between the 4th and 7th postfire sampling months. Production and shrub and sprout density increased substantially in the growing season. Results from the study are summarized below :
|Time since fire (months)||2||4||7||10|
|true mountain-mahogany plants/ha||0||10||5||40|
|Density of sprouts (#/ha)||0||153||79||1,453|
When individual true mountain-mahogany shrubs were burned with a kerosene flame gun, current annual growth increased with dormant season burning and decreased with growing season burning [326,327]. The degree to which this study imitated natural burning conditions is unknown. Curtin  reported that true mountain-mahogany was significantly (p-value not reported) lower on burned than unburned sites 2 years following wildfires in the Malpai Borderlands. Fire behavior was not described.
True mountain-mahogany was present on 1-year-old, 2-year-old, and unburned sites in the Eastwood manzanita (Arctostaphylos glandulosa) chaparral vegetation type in California's San Jacinto Mountains. Fires burned in late October and early September. The fire that burned 1-year-old sites consumed all litter and most shrub stems. The other fire was not described. No true mountain-mahogany seedlings occurred on burned or unburned sites. True mountain-mahogany cover and density were 0.1% and 40 individuals/acre on 1-year-old burned sites. Cover and density were not reported on 2-year-old sites but on unburned sites were 0.6% and 32 individuals/acre, respectively .
True mountain-mahogany frequency was the same on burned and unburned sites in a pinyon-Utah juniper community in west-central Utah. The area burned in August and nearly all of the nonsprouting vegetation was killed. True mountain-mahogany occurred in 1 of 16 plots on burned and unburned sites; the frequency was unchanged in the 1st, 2nd, and 3rd postfire years .
On Lower Covington Flats in Joshua Tree National Park, true mountain-mahogany frequency was lower but coverage was greater on burned than unburned sites. The fire burned in early July and denuded the valley floor but left some surviving plants on adjacent slopes. True mountain-mahogany occurred only on the slopes. Sites were visited 16 months following the fire, and some partially burned true mountain-mahogany survived. Frequency on burned and unburned sites was 9% and 12%, respectively. Coverage was 4.6% on unburned and 12% on burned sites .
Studies providing information on fire behavior: Typically mortality of true mountain-mahogany is greater on high-severity burned than low-severity burned sites. Survival of true mountain-mahogany stems was approximately 50% following a moderate to severe surface fire in blue oak woodlands of Sequoia National Park. The fire burned in late June when the temperature was around 86 °F (30 °C), relative humidity was 17%, and fine fuel moisture was 3.5%. Fire flame lengths were 3 to 15 feet (1-4.6 m). Some trees retained scorched leaves, while other trees had green upper crowns. Eighteen months after the fire, true mountain-mahogany had 3 live stems/ha and 3 dead stems/ha. The researcher indicated that "some" true mountain-mahogany shrubs sprouted . A wildfire that burned in Wyoming big sagebrush/true mountain-mahogany communities in south-central Wyoming caused an average of 25% true mountain-mahogany mortality. The fire burned in September when the temperature averaged 63 °F (17 °C), relative humidity averaged 33%, and wind speeds were 8.1 to 12 miles (13-20 km)/hour. True mountain-mahogany survival presented as a percentage of the prefire abundance was 82%, 78%, and 76% in the 1st, 2nd, and 3rd postfire years, respectively .
True mountain-mahogany cover and height on burned and unburned sites were not very different following underburning as part of the King River Ecosystems Project in the Sierra National Forest. Before the fires, true mountain-mahogany averaged 1.9% cover and 10.3 feet (3.14 m) in height. Postfire averages that spanned 2 to 8 years were 1.2% cover and 9.9 feet (3 m) in height .
Following December prescription fires in Jeffrey pine-blue oak vegetation in California's Cuyamaca Rancho State Park, true mountain-mahogany density was slightly greater on 1.5-year-old burned sites and slightly lower on 6-month-old burned sites than on control sites. Low-severity prescription fires burned under similar conditions. Temperatures ranged from 55 to 75 °F (13-24 °C), relative humidity was 17% to 37%, fuel moistures were 6% to 8%, and winds were 2.5 to 6 miles (4-9.6 km)/hour. On the 1.5-year-old burned site, true mountain-mahogany density was 3 plants/ha, and on control sites density was 0/ha. Density was 7 plants/ha and 9 plants/ha on 6-month-old burned and control sites, respectively .
True mountain-mahogany recovery was variable on a more severely burned site following the mid-June La Mesa Fire at Bandelier National Monument. Two sites dominated by ponderosa pine and oneseed juniper burned. Site 1 was unburned for 40 years prior to the fire. Fire spread across site 1 at 24 chains (66-1,584 feet)/hour. All trees had either brown needles or no needles remaining. In postfire year 1, total vegetation cover was 2.9%. Prefire true mountain-mahogany coverage was not reported. True mountain-mahogany was not recorded in postfire year 1 or 2, but coverage was 0.34% in postfire year 8 and 1.19% in postfire year 16. Site 2 had not burned for 17 years prior to the fire. Fire damage ranged from no scorch to 75% crown damage, and in postfire year 1 total vegetation cover was 9.4%. True mountain-mahogany prefire coverage was 3.4%, but true mountain-mahogany was not recorded 1, 2, 8, or 16 years after the fire .
Studies providing prefire and/or postfire site characteristics: Aspect, soils, and prefire stand characteristics were related to fire effects on true mountain-mahogany in the following studies. True mountain-mahogany coverage was greater on north than south slopes and on Andesite substrates than on sandstone substrates in early postfire shrublands in the Santa Monica mountains. The fire burned in the fall and recovery was evaluated approximately 6 and 7.5 months after the fire. Fire behavior characteristics were not described. Results of the study are summarized below .
|Time since fire (months)||6||7.5||6||7.5|
Pre- and postfire coverage of true mountain-mahogany was greater in younger Colorado pinyon-Utah juniper stands with lower canopy cover than in older stands in northeastern Utah's Daggett County. The behavior, season, or severity of the prescription fire were not described. Researchers recommended that a fire frequency that maintains Colorado pinyon-Utah juniper cover below 25% is best for the maintenance of true mountain-mahogany. Study findings are presented below :
|Approximate stand age (years)||80||300|
|Prefire Colorado pinyon-Utah juniper cover (%)||11.2||62.3|
|Prefire true mountain-mahogany cover (%)||11.4||1|
|True mountain-mahogany cover in postfire year 2||7.5||not measured|
|True mountain-mahogany cover in postfire year 4||9.5||0|
Postfire soils: Available nitrogen and extractable phosphorus increased by 14 and 4 times, respectively, after a fire in shrublands dominated by shrub live oak, true mountain-mahogany, and pointleaf manzanita in central Arizona's Prescott National Forest. Before the fire, available nitrogen in the soil under true mountain-mahogany shrubs was 5.4 mg/kg and phosphorus was 5.8 mg/kg. Relative humidity ranged from 10% to 30%, temperatures were 61 to 84 °F (16-29 °C), and wind speeds were 5 to 18 miles (8-29 km)/hour when the fire burned. True mountain-mahogany fuel moisture was 61% to 72% at the time of the fire. At its most "intense," fire flame heights were 10 feet (4 m), and fire spread was 39 feet (12 m)/minute. Approximately 70% of available fuels were consumed. Following the fire, soils under true mountain-mahogany had 71.6 mg/kg of available nitrogen and 22.6 mg/kg of available phosphorus . Another study of postfire soil nutrients in Arizona chaparral suggested that available nitrogen was unchanged and extractable phosphorus increased following a fall prescription fire. Researchers noted, though, that the distribution of available nitrogen and extractable phosphorus made it susceptible to erosion. Fire characteristics were very similar in the 2 studies .
Effects of repeated fire: Frequent fire may decrease true mountain-mahogany abundance. True mountain-mahogany had a frequency of 1.8% on sites burned once and 0.5% on sites burned twice in 2 years on Otay Mountain in San Diego County. On a west-facing slope outside the sampling area, true mountain-mahogany clump size was "severely" reduced, and some shrubs were killed following the 2nd fire. Within the perimeter of the 2nd fire were some small unburned patches. No other fire characteristics were provided .
Fire chronosequences: Studies reporting true mountain-mahogany abundance on past burned sites ranging from 4 to 400 years old are presented in an earlier section, see Postfire succession.
The Research Project Summary Response of vegetation to prescribed burning in a Jeffrey pine-California black oak woodland and a deergrass meadow at Cuyamaca State Park, California provides information on prescribed fire and postfire responses of many plant community species including birchleaf mountain-mahogany.FIRE MANAGEMENT CONSIDERATIONS:
Fuels and fire behavior: A prescribed fire study in chaparral of San Diego County showed that 45% of live and 95% of dead true mountain-mahogany fuels were consumed when burning occurred in the spring. Sites were treated with a desiccant to encourage burning. At the time of the fire, twigs that were 0.125 to 1 inch (0.3-2.5 cm) in diameter had 68% to 82% moisture, and fuels that were 2.5 to 3 inches (6.3-7.6 cm) in diameter had 47% moisture. Live fuels averaged 29 tons/acre, and dead fuels averaged 19 tons/acre. Minimum relative humidity was ~30%, maximum temperatures were 77 to 80 °F (25-27 °C), and winds were variable up to 10 miles (16 km)/hour with gusts of 12 to 15 mi (19-24 km)/hour when the fire burned. A fire temperature of 1,250 °F (677 °C) was recorded at 31 inches (79 cm) above the soil surface but likely does not represent a maximum temperature .
After a fire, true mountain-mahogany fuel structure is a large number of fairly uniform diameter sprouts, and as time without fire or aboveground destruction increases, stem mortality occurs, and eventually just a few large diameter stems persist. Based on the analysis of 9 shrubs between 4 and 18 years of age, 66% of woody fuels were less than 0.4 inch (1 cm) in diameter, and dead fuels were uncommon .
Prescription fire: Prescription and wildfire behavior are similar in chaparral vegetation; both produce crown fires. However, the severity of chaparral fires can vary. Low-, moderate-, and high-severity fires produce surface soil temperatures of ~480 °F (250 °C), 800 °F (425 °C), and 1,300 °F (700 °C), respectively. Temperatures at 1 inch (2.5 cm) depths are 210 °F (100 °C), 350 °F (175 °C), and 390 °F (200 °C) for low-, moderate-, and high-severity fires .
Green  provides information regarding the planning and execution of prescription fires in chaparral vegetation. However, the use of prescribed fire in chaparral vegetation is not considered necessary or useful by some. Keeley and Fotheringham  indicate that "there are few places where fire-dependent shrubs are threatened by a lack of fire" in southern California, and that the use of prescription fire as a means to mitigate future fire hazards is unwarranted. Prescribed burning is not likely to prevent fire spread under severe weather conditions, and expense would be great while resource benefit would be small. Zedler , too, suggests the prevention of fire through fire use is unrealistic and adds there is no "urgent need to reduce the time between fires" in southern California chaparral. He recommends that fire suppression efforts concentrate only on recently burned southern California shrublands, that controlled burning be discontinued, and that wildfires be allowed to burn, unless there is evidence of harm to an endangered species. He indicates that fires should be allowed to burn even in very old shrublands even under the "most severe circumstances" .
Postfire management: Heavy browsing of true mountain-mahogany postfire sprouts can kill shrubs. This warning came from observations made in Lake County, California .
Evaluating fire severity: Keeley and others  evaluated fire severity by measuring true mountain-mahogany postfire shrub heights in chaparral vegetation in the Sierra Nevada.
Dittberner and Olson  rate true mountain-mahogany forage as good for elk, fair to good for mule and white-tailed deer, poor for pronghorn, and fair for upland game birds, small nongame birds, and small mammals.
Livestock: True mountain-mahogany is rated good to excellent forage for all classes of livestock [53,78]. Forage is considered somewhat better for domestic sheep than cows .
Domestic goats in southern California consumed nearly 100% of the available browse of 1- and 5-year-old true mountain-mahogany shrubs. Researchers suggested that goats could be used to maintain fuel breaks to aid in the control of wildfires . When domestic goats are used to control juniper (Pinchot juniper and Ashe juniper (Juniperus ashei)) in Texas, heavy grazing of the area is required, and if true mountain-mahogany is present on browsed sites it will likely decrease since it is more palatable than juniper .
Response of true mountain-mahogany to browsing is provided in the Response to browsing section below.
Livestock and wildlife: Utilization of true mountain-mahogany by both wildlife and livestock requires that habitats be managed to avoid over use. True mountain-mahogany may be consumed by both livestock and wildlife species any time these species occupy the same area. Mule deer, cattle, and domestic sheep utilized 70% of leaves and annual stem growth of true mountain-mahogany on "properly" stocked areas on northern Arizona's Kaibab National Forest . True mountain-mahogany is referred to as a "superior feed" in California . True mountain-mahogany/sideoats grama habitat type in the Black Hills National Forest provides late spring and early summer livestock range and big game winter range. Researchers indicate that proper grazing management is essential to maintain deer winter range . Colorado pinyon-alligator juniper/true mountain-mahogany-gray oak/mountain muhly vegetation in western New Mexico has high early-season grazing potential and receives moderate big game use . On south-central Utah's Oak Creek range, true mountain-mahogany utilization was high in areas where either deer or cattle sign were plentiful. Use of true mountain-mahogany areas available only to deer was 34.4%, but on areas used by both deer and cattle, use was 53.6%. Areas with both deer and cattle averaged 1.7 times as many deer as did areas used by deer alone .
When true mountain-mahogany grows into a small tree form, much of its available browse is out of the reach of livestock and big game animals . Following fire, heavy browsing of new true mountain-mahogany sprouts can kill shrubs .
Deer: Mule and white-tailed deer consume true mountain-mahogany and utilize true mountain-mahogany habitats year round throughout its range [39,78,209,210]. Degree of use may depend on phenology, season, and/or weather.
True mountain-mahogany is typically rated as highly valuable or excellent forage [39,78,209,210]. Mule deer diet information that was summarized from studies in the western United States indicated that true mountain-mahogany received moderate use in the spring and fall and heavy use in the winter and summer based on 8 and 15 studies, respectively .
Contents of 96 mule deer stomachs taken from the Tehama winter range in northern California revealed that consumption of true mountain-mahogany was greatest in April and May. Usage was lowest in the fall and winter. The frequency of true mountain-mahogany in mule deer stomachs was 28.5% for this time . Based on 328 fecal samples collected in southeastern Wyoming's Medicine Bow National Forest, the frequency of true mountain-mahogany in mule deer diets was 13% in the summer, 8% in the fall, 50% in the winter, and 46% in the spring. Winter and spring samples were collected in an area dominated by true mountain-mahogany . In central Arizona's Sierra Ancha Experimental Forest, utilization by mule and white-tailed deer was greatest in the spring, 36%, and low to none until November when utilization was 13% .
Monitoring data from 22 radio-collared adult female mule deer indicated that true mountain-mahogany stands were "preferred" for feeding and for resting in the daylight hours. Monitoring was done from mid-November through March for 3 years in Colorado's Front Range . Based on the volume of true mountain-mahogany in mule and white-tailed deer stomachs and its availability in the Three Bar Wildlife Area of south-central Arizona, researchers determined that true mountain-mahogany was not necessarily a preferred food source. Utilization did, however, increase in late winter . In Guadalupe National Park in Texas, researchers determined that the relative density of true mountain-mahogany was 13% and frequency was 74% in annual mule deer diets. Fecal and rumen analysis and observations were used to assess consumption .
In the eastern part of Carlsbad Caverns National Park, mule deer fed on true mountain-mahogany for 7 of the 222.5 minutes of observations in a "good growing season" and in 6 of the total 158 observations. Feeding on true mountain-mahogany was not observed in a "poor growing season." In the Park's Scoggins Corner area, the frequency of true mountain-mahogany in 6 deer analyzed in a "good growing season" was 33% and in a poor growing season from 12 deer was 17% . True mountain-mahogany made up only 0.01% of the vegetative cover on New Mexico's Fort Stanton Experimental Ranch, but percent composition of mule deer diets in the area as determined through fecal analyses were 6.9%, 5.0%, and 5.8% in June, September, and March, respectively .
Several studies that involved the use of tame deer were not included in the above discussion. If interested, see [21,48,201].
Management issues related to deer use: Heavy winter browsing of true mountain-mahogany in California has caused severe shrub damage . Chaparral vegetation on the San Joaquin deer winter range in Madera County, California, that was too tall or dense to provide deer browse was smashed by a bull dozer and then burned in early spring. After 6 years of intense deer browsing (utilization of 35% to 85%) of postfire sprouts, only a few grew out of deer's reach. On 22-year-old burned and browsed sites, true mountain-mahogany grew into a tree-like form largely beyond the reach of deer. Even with prolonged intense use, sprouts can become unavailable after time [80,81]. For more information on true mountain-mahogany and its response to browsing pressure, see Response to browsing.
Deer and sheep: In Palo Duri Canyon, Texas, researchers found a high degree of overlap in the diets of Barbary sheep and mule deer. The relative density of true mountain-mahogany was 12.5% and 11.9% in Barbary sheep and mule deer diets, respectively, based of the analysis of feces collected from February 1977 to January 1979. Sand shinnery oak (Quercus havardii) made up bulk of both species diets and contributed most to diet overlap .
Bighorn sheep: True mountain-mahogany is consumed primarily in the summer by bighorn sheep. True mountain-mahogany made up the greatest content in 3 bighorn sheep stomachs taken from Texas in the early 1900s (Bailey in ). No true mountain-mahogany was recovered from bighorn sheep feces collected in the fall or winter in the Sheep Creek-Trickle Mountain Area of Saguache County, Colorado. Bighorn sheep did feed on true mountain-mahogany in the summer and spring. True mountain-mahogany made up a trace of spring diets and 12.1% of the relative density of feces collected in the summer .
Bighorn sheep in Waterton Canyon, Colorado, preferred mountain shrub habitats between east and southwest aspects. True mountain-mahogany was the dominant shrub on south slopes . Based on analysis of collected feces and direct feeding observations, researchers found that true mountain-mahogany leaves dominated the summer diet of bighorn sheep. From fecal analyses, true mountain-mahogany made up 74% to 90% of the relative densities of diets from June through September. Observations generally agreed with fecal analyses findings .
Deer and elk: True mountain-mahogany is thought to be consumed more heavily by deer than elk , but both game animals utilize true mountain-mahogany habitats and browse. However, season of consumption may differ. Colorado pinyon/true mountain-mahogany/nodding brome vegetation in western New Mexico is considered excellent deer habitat but receives elk use, too . Usage of Colorado pinyon-alligator juniper vegetation where true mountain-mahogany was important, as determined by deer and elk pellet counts, increased as shrub coverage increased in the Gila National Forest of southern New Mexico .
Researchers compared the diets of mule deer and elk in Colorado pinyon-oneseed juniper and ponderosa pine woodlands in north-central New Mexico. Based on year-round feces collections that spanned 2 years, true mountain-mahogany was 5% and 16% of annual elk and mule deer diets, respectively. Utilization of true mountain-mahogany was greatest in the summer for elk when it made up 12% the diets, and mule deer diets were 23% to 24% true mountain-mahogany in the summer and fall . Both elk and mule deer fed heavily on true mountain-mahogany in the Fort Baynard Watershed in New Mexico, but season of heaviest use differed. True mountain-mahogany made up 17% to 39% of 43 elk rumens. The low of 17% was found in the winter and the high of 39% was in the spring. For mule deer, stomachs contained 15% to 45% true mountain-mahogany. The highest amount of true mountain-mahogany was from collections made in the fall and winter, and the lowest was from summer collections .
Neither deer nor elk utilized true mountain-mahogany from January through March on northern Utah winter ranges. Based on fecal analyses, true mountain-mahogany was absent from mule deer diets and was less than 1% of the relative density of elk diets .
Elk: Based on feeding observations in Wind Cave National Park in the northern Great Plains, elk use of true mountain-mahogany was greatest in the spring. Fall, winter, spring, and summer use was 0%, 3.4%, 6.8%, and 1.1%, respectively .
Pronghorn: Researchers have observed pronghorn feeding on true mountain-mahogany and detected true mountain-mahogany in pronghorn feces. In north-central Colorado, researchers observed pronghorn feeding on true mountain-mahogany, but percentage of the diet or preference were not discussed . In Utah the maximum percentage of true mountain-mahogany in pronghorn diets was 15% based on rumen or feces collected in the spring or summer .
Coyotes: In Las Animas County, Colorado, researchers found that 72 radio-collared coyotes used canyon shrub-grassland communities dominated by skunkbush sumac and true mountain-mahogany more than expected (p<0.005) based on habitat availability. Of 18 occupied dens, 7 were in the shrub-grassland type .
Collared peccaries: Researchers found that true mountain-mahogany was one of the most common shrubs found at javelina bed sites in urban areas of Prescott, Arizona. Bed sites were located by tracking 8 collared peccaries from 6 different herds .
Small mammals: Many small mammals feed on true mountain-mahogany and utilize its habitats. In the Platte River Watershed in Colorado, prairie voles and deer mice were captured most often in the true mountain-mahogany-dominated vegetation type . Brush mice utilized true mountain-mahogany-dominated vegetation in the Granite Basin area of Arizona's Yavapai County. Nearly 30% of all trappings occurred in true mountain-mahogany-dominated stands. True mountain-mahogany stands averaged 4.3 captures/trap night .
In California's San Gabriel Mountains, steep rocky slopes dominated by true mountain-mahogany and Nuttall's scrub oak were habitat for brush mouse, California mouse, and dusky-footed woodrat populations. In a single acre of Nuttall's scrub oak-true mountain-mahogany shrublands near Jackson Lake, there were 13 occupied dusky-footed woodrat nests . Researchers examined the food contents in dusky-footed woodrat nests on the San Dimas Experimental Forest in Los Angeles County, California. The frequency of true mountain-mahogany leaves in nests was as high as 100% on some sites, and the maximum distance leaves were carried was 45 feet (14 m). True mountain-mahogany is an important summer food. Over 0.02 pound (7 g) of true mountain-mahogany was recovered from nests in July and September, but true mountain-mahogany occurred in nests in all seasons .
True mountain-mahogany is typical in oak chaparral and oak-juniper habitats in Brewster County, Texas, utilized by white-ankled mice, Merriam's pocket mice, brush mice, cactus mice, yellow-nosed cotton rats, western white-throated woodrats, desert cottontails, black-tailed jackrabbits, rock squirrels, Texas antelope squirrels, raccoons, ringtails, and gray foxes. Specific use of true mountain-mahogany was not discussed .
Based on fecal analyses made from summer collections from Colorado pinyon-Utah juniper woodlands of Colorado's Piceance Basin, researchers found that true mountain-mahogany made up 1.1% and 2.2% of the relative density of deer mouse and least chipmunk habitats, respectively .
Birds: True mountain-mahogany provides cover and habitat to many game and song birds. Scaled quail in Oldham County, Texas, used true mountain-mahogany habitats in the summer . One of seven located Montezuma quail nests in the Huachuca Mountains of Cochise County, Arizona, occurred on hot, dry limestone ridges where cover included true mountain-mahogany . Seventy night time scaled quail roost sites were found in the Rolling Plains of northwestern Texas in the spring and winter. The true mountain-mahogany-yucca (Yucca spp.)-fragrant sumac vegetation type was selected by scaled quail based on calculated electivity ratios. Eighteen of the located roosts were in this vegetation type .
Between 19 and 32 species of breeding birds occurred in upland forests of east-central Taos County, New Mexico. True mountain-mahogany was a common shrub in these forests. However, specific use of true mountain-mahogany was not mentioned . There were 9 species of breeding birds and a total of 103 breeding pairs/40 ha in shrub live oak-true mountain-mahogany chaparral vegetation in central Arizona's Tonto National Forest. The following bird species occurred in a greater numbers in shrub live oak-true mountain-mahogany chaparral vegetation than in nearby grassland or riparian sites: Gambel's quail, common poorwill, bridled titmouse, Bewick's wren, crissal thrasher, northern cardinal, and eastern towhee .
True mountain-mahogany was common in Virginia's warbler breeding habitats in the southwestern Black Hills of South Dakota. Researchers often observed Virginia's warblers foraging in true mountain-mahogany shrubs. It is likely that true mountain-mahogany was important nest cover, too. The density of breeding Virginia's warblers was 8.3 birds/km² . In the Sandia Mountains of north-central New Mexico, a crissal thrasher nest was located 2 feet (0.6 m) above the ground in a true mountain-mahogany shrub in Embudo Canyon .
Palatability/nutritional value: True mountain-mahogany is considered a "highly preferred," "highly palatable" browse species [123,202]. Stems are considered palatable year round . Palatability rates excellent to very good for deer, bighorn sheep, and domestic goats, good to very good for cattle and goats, and good to poor for horses [128,245,271]. True mountain-mahogany is considered an excellent source of nitrogen for livestock and big game animals . Forage collected in the winter near Logan, Utah, was fed to captive deer. Based on browsing time and forage consumed, true mountain-mahogany × curlleaf mountain-mahogany hybrids were considered more palatable than true mountain-mahogany species .
True mountain-mahogany in vitro digestibility, crude protein, and phosphorus average 26.5%, 7.8%, and 0.13%, respectively, in the winter season. These averages are based on values reported in the literature. Given these values, a diet of only true mountain-mahogany is deficient in meeting crude protein, total digestible nutrients, and phosphorus requirements of wintering mule deer but exceeds calcium and carotene requirements . Based on the nutrient requirements for cattle, domestic sheep, horses, deer, elk, small mammals, and birds, a diet of only true mountain-mahogany in the winter would be deficient for all but cattle during gestation, lactation, growth, and maintenance periods. For cattle, a diet of only true mountain-mahogany would be adequate only during maintenance and gestation periods . In the Uinta Mountains of Utah, researchers reported nearly toxic levels of copper in true mountain-mahogany. Poisoning would depend on the amount of true mountain-mahogany consumed .
The current year's growth of mountain-mahogany (Cercocarpus spp.) leaves and stems were higher in digestible organic matter and crude protein only in the 1st postfire growing season when browse on control, 1-, 2-, 3-, and 6-year-old burned sites was compared in Guadalupe Mountains of New Mexico .
Many studies have investigated the nutritional content of true mountain-mahogany leaves and stems. Some studies analyzed stems and leaves separately while other did not discriminate. Some were interested in seasonal fluctuations while others were season specific. Some report that crude protein levels do not fluctuate seasonally [22,294], while others indicate that crude protein decreases from spring to winter . Refer to the studies listed below for information on true mountain-mahogany nutritional and/or chemical content.
|Study area||Plant portion collected||Season of collection||Notes||Reference|
|Arizona||current year's growth; stems and leaves analyzed separately||late September, late November, early January, early March||provides in vitro digestibility for mule and white-tailed deer separately|||
|Arizona, Mogollon Plateau||leaves||June, August|||
|Arizona, Sierra Ancha Experimental Forest||terminal 2 inches (5 cm) of sprout growth||year round|||
|California||current year's growth stems and leaves||year round||crude protein only|||
|California, North Coast Range||current year's growth||year round||comparisons for 2-, 3-, and 5-year-old burned sites|||
|Colorado, western||current year's growth||January|||
|New Mexico, north-central in Jemez Mountains||current year's growth||January-April||3-year-old burned site|||
|New Mexico, southwest in Fort Baynard Watershed||current year's growth||year round|||
|Wyoming||current year's growth||growing season|||
|Utah, central||leaves and stems analyzed separately||April, July, September, November||collections in April were before leaf out; in July relatively little growth; in November temperatures freezing|||
|Utah, Uinta Basin||current year's growth; stem base, middle, and tip analyzed separately||early October|||
|Utah, Uinta Mountains||stems within easy reach of large herbivores||early October|||
True mountain-mahogany likely provides cover for many wildlife species. The oak chaparral vegetation of
Brewster County, Texas, where true mountain-mahogany is common, provides good cover for black-tailed
jackrabbits and desert cottontails . True mountain-mahogany provides excellent hiding or escape cover
and fawning cover and good thermal cover for mule deer in Wyoming . True mountain-mahogany is considered
fair hiding/escape and thermal cover and good fawning cover for white-tailed deer in Wyoming . Dittberner
and Olson  rate true mountain-mahogany cover as good for mule and white-tailed deer, fair for elk, poor for
pronghorn, and fair to good for upland game birds, small nongame birds, and small mammals.
VALUE FOR REHABILITATION OF DISTURBED SITES:
Many consider true mountain-mahogany a valuable component in revegetation or reclamation efforts. Seed and/or plants are available commercially [118,291]. True mountain-mahogany provides soil stabilization [125,179], has fair to good drought tolerance , fixes nitrogen [3,322], and provides important wildlife browse and cover .
Site conditions and community types within the Intermountain West that are best suited for true mountain-mahogany revegetation are provided in . Guidelines on how and where to plant true mountain-mahogany in New Mexico are given in . True mountain-mahogany seed collection, cleaning, and storage information is available in , and guidelines for producing true mountain-mahogany container seedlings is summarized in .
True mountain-mahogany is difficult to establish from seed , and low seedling "vigor" and "slow initial growth rates" are typical . Seed may be locally adapted and may not do well without proper stratification. Using seed collected near the proposed planting site is recommended [152,195]. Seeding is typically done in the fall; for information on seedbed preparation and seeding rates, see . True mountain-mahogany seed needs to be planted in the soil at depth of ~0.5 inch (1 cm); broadcast seeding is seldom successful. Researchers recommend against planting true mountain-mahogany with herbaceous species. For more seeding and/or planting tips, see . If planting conditions are favorable, establishment and survival are not compromised by low initial growth rates . Successful establishment of true mountain-mahogany from bareroot stock is considered moderate. Container-grown plants do not establish as well . Due to the slow establishment of true mountain-mahogany (4-6 years), revegetated sites should be protected from livestock grazing for at least 4 years, and protection from native browsers may also be necessary .
Results of revegetation projects: Researchers seeded and planted container grown true mountain-mahogany on 40-year-old burned and unburned open ponderosa pine stands in South Dakota's Black Hills National Forest. Plantings were protected from browsers and monitored for 10 years. Seeded true mountain-mahogany survival averaged 4.2% on burned and 3.6% on unburned open stands in postfire year 10. Container grown plantings survived better; survival averaged 6.7% on burned and 18.2% in unburned open stands . Survival of true mountain-mahogany was as high as 95% six years after planting container-grown plants in southwestern Utah .
Results of mine reclamation projects:
True mountain-mahogany survival was 25% two years after planting on a surface-mine site in Oak Creek,
Colorado. Survival did not differ on overburden sites and sites with topsoil . True mountain-mahogany
survival was 0% on surface coal mined reclamation sites in grassland vegetation of Colorado but was 25% on
mine sites in mixed shrub vegetation sites .
Native people utilized true mountain-mahogany in the past. Wood was used by native people of southern California in making arrow tips, digging tools, spears, and fighting clubs [41,50]. Navajo people used true mountain-mahogany to make their weaving tools and other tool handles. True mountain-mahogany wood was used in the construction of sweat houses and prayer sticks utilized in the Mountain Chant Ceremony [66,299]. Hopi people used true mountain-mahogany bark to make a red-brown dye for leather . This type of dye was also utilized by the Ramah Navaho of northwestern New Mexico to color buckskin, wool, and basket materials . Kawaisu natives of south-central California carved pipe heads from true mountain-mahogany .
True mountain-mahogany was also used medicinally. Navajo people took true mountain-mahogany extracts to treat stomach problems, such as sickness from overeating venison, and to speed up postpartum recovery. Chewing true mountain-mahogany leaves from a deer-browsed plant was thought to bring a Navajo hunter good luck [66,299]. Native people of southeastern Utah reportedly utilized true mountain-mahogany as a medicine but what it treated is unknown . Kawaisu natives of south-central California boiled the root of true mountain-mahogany and drank it to treat coughing .
More recently true mountain-mahogany has been suggested as a landscape or garden plant in the northwestern and southwestern U.S. and in the southern Great Plains [95,126,157]. It is suggested for dry, well-drained sites  and has been recommended for water-wise landscaping because of its drought and heat tolerance .
True mountain-mahogany may also have a future use. Arizona chaparral species including true mountain-mahogany showed promise as "feedstocks for pyrolytic conversion to liquid fuels." For more information, see .
Wood produced by true mountain-mahogany is hard and brittle
OTHER MANAGEMENT CONSIDERATIONS:
Response to browsing: While true mountain-mahogany tolerates "close" browsing , it can be killed by heavy use . It has been referred to as both a decreaser and increaser in the literature. Parker  considers true mountain-mahogany a "decreaser" when browsed yearlong by domestic sheep, cattle, and deer but refers to true mountain-mahogany as an "increaser" on upland or mountain sites when only cattle are the nonnative browsers. Zacek and others  indicate that true mountain-mahogany decreases in relative abundance with continued heavy use. Likely site conditions, grazing timing, and climatic stress all play a role in the browsing response.
True mountain-mahogany cover was lower after 18 years of light and moderate use by cattle in Colorado's Manitou Experimental Forest. Before cattle were on the sites, true mountain-mahogany cover was 0.12% and 0.05% on lightly and moderately used sites, respectively. After the introduction of cattle, coverage was 0.05% on lightly used and 0.02% on moderately used sites . In Rio Blanco County, Colorado, 3 relatively homogeneous enclosures were utilized at different levels by different herbivores. Enclosure 1 supported 2 domestic sheep/acre/month during the spring and fall and 2.3 deer/acre/month in the winter. Enclosure 2 was utilized at 15.4 cows/acre/month in the spring and fall and 2.3 deer/acre/month in the winter. Enclosure 3 had 1.6 deer/acre/month in the winter. Percent composition of true mountain-mahogany was lowest in enclosure 1, and sheep browsing had decreased plant "vigor" and killed some plants. Composition or frequency of true mountain-mahogany in enclosures 2 and 3 was not different, but cattle and deer browsing on true mountain-mahogany was heavy. Heavy use of true mountain-mahogany by sheep may decrease browse available to deer using the same area .
Researchers compared true mountain-mahogany growth inside and outside an exclosure in the Medicine Bow National Forest of Wyoming. Vegetation within the exclosure was protected for 40 years. True mountain-mahogany density and canopy cover were similar inside and outside of the exclosure. Production was significantly (p=0.1) higher outside (242.3 kg/ha) than inside (107.8 kg/ha), and the amount of dead true mountain-mahogany material was significantly greater inside (43.5%) than outside (18.7%). Researchers suggested that some level of browsing may be necessary to maintain an "optimum sustained yield" of true mountain-mahogany .
Browsed and unbrowsed true mountain-mahogany shrubs in the Wasatch and Uinta mountains of Utah were more affected by resource availability than by previous year's use. Regardless of use, annual twig growth and flower and seed production were greatest when precipitation levels were greatest. Utilization of shrubs ranged from <21% to over 300% (consumption of annual growth and previous year's growth). Shrubs compensated but did not over compensate for losses; browsed shrubs were smaller than unbrowsed shrubs. Flower and seed production were much greater on unbrowsed than browsed shrubs. Herbivores in the area were moose, elk, deer, antelope, and cattle [288,289].
Browsing simulated by clipping: Clipped shrubs produced more biomass than unclipped shrubs in Logan, Utah. Ninety percent of the previous year's growth of 4-year-old shrubs was clipped in either December or April. When 90% of the distal buds were removed in the spring, true mountain-mahogany produced significantly (p<0.05) more biomass than control plants. True mountain-mahogany also averaged 1 new sprout/plant following the treatment. Experiments were conducted in manipulated environments where nutrients and light availability were high, unlike natural conditions .
Season of clipping affected the response of true mountain-mahogany in Ft. Collins, Colorado. Researchers removed 80% and 40% of stems at ~2 inches (5 cm) above the ground. Current annual growth of clipped plants was higher than controls when plants were clipped in the dormant season and lower when plants were clipped during the growing season. Differences, however, were not significant . Twelve years of clipping true mountain-mahogany in southwestern Colorado suggests that shrubs are resistant to browsing levels of 60% to 80%. No significant differences were found between the current annual growth or mean stem length of control shrubs and shrubs with 20% to 100% of their current annual growth removed. Plants receiving 100% removal treatments were monitored for 10 years. Seed production was lowest for heavily clipped shrubs. Shrubs that had all current annual growth removed and old wood to 0.2 inch (4 mm) removed produced new sprouts early in the treatments, but none survived over 6 years of this treatment .
Predictive equations: Predictive equations for estimating true mountain-mahogany fuels in southern California chaparral are provided in . Brotherson and others  used diameter and age data to develop predictive equations for populations in the Laramie Basin of Wyoming and the Uinta Mountains of Utah. Combined data from both populations produced more accurate estimates for either population, and researchers suggested that similar true mountain-mahogany growth rates made equations useful to both areas. Researchers used twig length and diameter measurements of true mountain-mahogany current year's growth, including leaves, in a regression equation to predict production and utilization. Measurements were obtained from shrubs in the Organ Mountains Recreation Lands in southern New Mexico. Predictive power for other populations was not assessed .
Methods for increasing production: Dimeo  provides a review of the mechanical tools used (successfully and unsuccessfully) to increase true mountain-mahogany browse.
Methods to control: The control of postfire true mountain-mahogany sprouts as a means to increase herbaceous species production in chaparral vegetation in San Gabriel Mountains is described in .
1. Ackerly, David. 2004. Functional strategies of chaparral shrubs in relation to seasonal water deficit and disturbance. Ecological Monographs. 74(1): 25-44. 
2. Agee, James K.; Biswell, Harold H. 1978. The fire management plan for Pinnacles National Monument. In: Linn, Robert M., ed. Proceedings, 1st conference on scientific research in the National Parks: Vol. 2; 1976 November 9-12; New Orleans, LA. NPS Transactions and Proceedings No. 5. Washington, DC: U.S. Department of the Interior, National Park Service: 1231-1238. 
3. Aldon, Earl F. 1978. Endomycorrhizae enhance shrub growth and survival on mine spoils. In: Wright, Robert A., ed. The reclamation of disturbed arid lands. Albuquerque, NM: University of New Mexico Press: 174-179. 
4. Allen, Barbara H.; Holzman, Barbara A.; Evett, Rand R. 1991. A classification system for California's hardwood rangelands. Hilgardia. 59(2): 1-45. 
5. Allen-Diaz, Barbara H.; Holzman, Barbara A. 1991. Blue oak communities in California. Madrono. 38(2): 80-95. 
6. Allison, Chris. 1988. Seeding New Mexico rangeland. Circular 525. Las Cruces, NM: New Mexico State University, College of Agriculture and Home Economics, Cooperative Extension Service. 15 p. 
7. Andersen, Berniece A.; Holmgren, Arthur H. . Mountain plants of northeastern Utah. Circular 319. Logan, UT: Utah State University, Extension Services. 148 p. 
8. Anderson, David Lee. 1974. Ecological aspects of Cercocarpus montanus Raf. communities in central Utah. Provo, UT: Brigham Young University. 84 p. Thesis. 
9. Arno, Stephen F. 1980. Forest fire history in the Northern Rockies. Journal of Forestry. 78(8): 460-465. 
10. Arno, Stephen F. 2000. Fire in western forest ecosystems. In: Brown, James K.; Smith, Jane Kapler, eds. Wildland fire in ecosystems: Effects of fire on flora. Gen. Tech. Rep. RMRS-GTR-42-vol. 2. Ogden, UT: U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station: 97-120. 
11. Arno, Stephen F.; Gruell, George E. 1983. Fire history at the forest-grassland ecotone in southwestern Montana. Journal of Range Management. 36(3): 332-336. 
12. Arno, Stephen F.; Scott, Joe H.; Hartwell, Michael G. 1995. Age-class structure of old growth ponderosa pine/Douglas-fir stands and its relationship to fire history. Res. Pap. INT-RP-481. Ogden, UT: U.S. Department of Agriculture, Forest Service, Intermountain Research Station. 25 p. 
13. Arno, Stephen F.; Wilson, Andrew E. 1986. Dating past fires in curlleaf mountain-mahogany communities. Journal of Range Management. 39(3): 241-243. 
14. Arthun, Dave; Holechek, J. L.; Wallace, J. D.; Gaylean, M. L.; Cardenas, M. 1992. Forb and shrub effects on ruminal fermentation in cattle. Journal of Range Management. 45: 519-522. 
15. Baisan, Christopher H.; Swetnam, Thomas W. 1990. Fire history on a desert mountain range: Rincon Mountain Wilderness, Arizona, U.S.A. Canadian Journal of Forest Research. 20: 1559-1569. 
16. Baker, F. S.; Korstian, Clarence F. 1931. Suitability of brush lands in the Intermountain region for the growth of natural or planted western yellow pine forests. Tech. Bull. No. 256. Washington, DC: U.S. Department of Agriculture. 82 p. 
17. Baker, William L. 1984. A preliminary classification of the natural vegetation of Colorado. The Great Basin Naturalist. 44(4): 647-676. 
18. Baker, William L.; Kennedy, Susan C. 1985. Presettlement vegetation of part of northwestern Moffat County, Colorado, described from remnants. The Great Basin Naturalist. 45(4): 747-783. 
19. Barrett, Stephen W. 1993. Fire regimes on the Clearwater and Nez Perce National Forests north-central Idaho. Final Report: Order No. 43-0276-3-0112. Ogden, UT: U.S. Department of Agriculture, Forest Service, Intermountain Research Station, Fire Sciences Laboratory. Unpublished report on file with: U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory, Missoula, MT. 21 p. 
20. Barrett, Stephen W.; Arno, Stephen F.; Key, Carl H. 1991. Fire regimes of western larch - lodgepole pine forests in Glacier National Park, Montana. Canadian Journal of Forest Research. 21: 1711-1720. 
21. Bartmann, Richard M. 1983. Composition and quality of mule deer diets on pinyon-juniper winter range, Colorado. Journal of Range Management. 36(4): 534-541. 
22. Bedell, Thomas E. 1980. Range nutrition in relation to management. Extension Circular 1045. Corvallis, OR: Oregon State University, Extension Service. 4 p. 
23. Bernard, Stephen R.; Brown, Kenneth F. 1977. Distribution of mammals, reptiles, and amphibians by BLM physiographic regions and A.W. Kuchler's associations for the eleven western states. Tech. Note 301. Denver, CO: U.S. Department of the Interior, Bureau of Land Management. 169 p. 
24. Berndt, Herbert W.; Gibbons, Robert D. 1958. Root distribution of some native trees and understory plants growing on three sites within ponderosa pine watersheds in Colorado. Station Paper No. 37. Fort Collins, CO: U.S. Department of Agriculture, Forest Service, Rocky Mountain Forest and Range Experiment Station. 14 p. 
25. Bissell, Harold D.; Strong, Helen. 1955. The crude protein variations in the browse diet of California deer. California Fish and Game. 41(2): 145-155. 
26. Biswell, H. H. 1961. Manipulation of chamise brush for deer range improvement. California Fish and Game. 47(2): 125-144. 
27. Biswell, H. H.; Gilman, J. H. 1961. Brush management in relation to fire and other environmental factors on the Tehama deer winter range. California Fish and Game. 47(4): 357-389. 
28. Biswell, H. H.; Schultz, A. M.; Hedrick, D. W. 1953. A possible method of increasing western mountain mahogany on game ranges. California Fish and Game. 39(2): 187-189. 
29. Booth, D. Terrance; Schuman, Gerald E. 1981. Shrub reestablishment research at the High Plains Grasslands Research Station. In: Stelter, Lavern H.; DePuit, Edward J.; Mikol, Sharon A., tech. coords. Shrub establishment on disturbed arid and semi-arid lands: Proceedings of the symposium; 1980 December 2-3; Laramie, WY. Cheyenne, WY: Wyoming Game and Fish Department: 81-88. 
30. Booth, W. E.; Wright, J. C. 1962. [Revised]. Flora of Montana: Part II--Dicotyledons. Bozeman, MT: Montana State College, Department of Botany and Bacteriology. 280 p. 
31. Borchert, Mark I.; Cunha, Nancy D.; Krosse, Patricia C.; Lawrence, Marcee L. 1993. Blue oak plant communities of southern San Luis Obispo and northern Santa Barbara Counties, California. Gen. Tech. Rep. PSW-GTR-139. Albany, CA: U.S. Department of Agriculture, Forest Service, Pacific Southwest Research Station. 49 p. 
32. Boyd, Steve. 1999. Vascular flora of the Liebre Mountains, western Transverse Ranges, California. Aliso. 18(2): 93-139. 
33. Bradley, Anne F.; Noste, Nonan V.; Fischer, William C. 1991. Fire ecology of forests and woodlands in Utah. Gen. Tech. Rep. INT-287. Ogden, UT: U.S. Department of Agriculture, Forest Service, Intermountain Research Station. 128 p. 
34. Brooks, Asa C. 1962. An ecological study of Cercocarpus montanus and adjacent communities in part of the Laramie Basin. Laramie, WY: University of Wyoming. 53 p. Thesis. 
35. Brotherson, Jack D. 1990. Mineral-nutrient relationships of the mountain mahoganies (Cercocarpus). In: Johnson, Kendall L., ed. Proceedings, 5th Utah shrub ecology workshop: The genus Cercocarpus; 1988 July 13-14; Logan, UT. Logan, UT: Utah State University, College of Natural Resources: 43-60. 
36. Brotherson, Jack D.; Anderson, D. L.; Szyska, L. A. 1984. Habitat relations of Cercocarpus montanus (true mountain mahogany) in central Utah. Journal of Range Management. 37(4): 321-324. 
37. Brotherson, Jack D.; Davis, James N.; Greenwood, Larry. 1980. Diameter-age relationships of two species of mountain mahogany. Journal of Range Management. 33(5): 367-370. 
38. Brotherson, Jack D.; Osayande, Solomon T. 1980. Mineral concentrations in true mountain mahogany and Utah juniper, and in associated soils. Journal of Range Management. 33(3): 182-185. 
39. Bryant, Fred C.; Demarais, Steve. 1991. Habitat management guidelines for white-tailed deer in south and west Texas. In: Lutz, R. Scott; Wester, David B., editors. Research highlights--1991: Noxious brush and weed control; range and wildlife management. Volume 22. Lubbock, TX: Texas Tech University, College of Agricultural Sciences: 9-13. 
40. Carmichael, R. S.; Knipe, O. D.; Pase, C. P.; Brady, W. W. 1978. Arizona chaparral: plant associations and ecology. Res. Pap. RM-202. Fort Collins, CO: U.S. Department of Agriculture, Forest Service, Rocky Mountain Forest and Range Experiment Station. 16 p. 
41. Chesnut, V. K. 1902. Plants used by the Indians of Mendocino County, California. Contributions from the U.S. National Herbarium. [Washington, DC]: U.S. Department of Agriculture, Division of Botany. 7(3): 295-408. 
42. Collins, Ellen I. 1984. Preliminary classification of Wyoming plant communities. Cheyenne, WY: Wyoming Natural Heritage Program/The Nature Conservancy. 42 p. 
43. Conrad, C. Eugene. 1987. Common shrubs of chaparral and associated ecosystems of southern California. Gen. Tech. Rep. PSW-99. Berkeley, CA: U.S. Department of Agriculture, Forest Service, Pacific Southwest Forest and Range Experiment Station. 86 p. 
44. Cook, John G.; Hershey, Terry J.; Irwin, Larry L. 1994. Vegetative response to burning on Wyoming mountain-shrub big game ranges. Journal of Range Management. 47(4): 296-302. 
45. Cooper, Charles F. 1960. Changes in vegetation, structure, and growth of southwestern pine forests since white settlement. Ecological Monographs. 30(2): 129-164. 
46. Costello, David F.; Price, Raymond. 1939. Weather and plant-development data as determinants of grazing periods on mountain range. Tech. Bull. 686. Washington, DC: U.S. Department of Agriculture. 31 p. 
47. Cronquist, Arthur; Holmgren, Noel H.; Holmgren, Patricia K. 1997. Intermountain flora: Vascular plants of the Intermountain West, U.S.A. Vol. 3, Part A: Subclass Rosidae (except Fabales). New York: The New York Botanical Garden. 446 p. 
48. Currie, P. O.; Reichert, D. W.; Malechek, J. C.; Wallmo, O. C. 1977. Forage selection comparisons for mule deer and cattle under managed ponderosa pine. Journal of Range Management. 30(5): 352-356. 
49. Curtin, Charles G. 2003. Fire as a landscape restoration and management tool in the Malpai Borderlands. In: Galley, Krista E. M.; Klinger, Robert C.; Sugihara, Neil G., eds. Proceedings of fire conference 2000: the first national congress on fire ecology, prevention, and management; 2000 November 27-December 1; San Diego, CA. Miscellaneous Publication No. 13. Tallahassee, FL: Tall Timbers Research Station: 79-87. 
50. Dale, Nancy. 1986. Flowering plants: The Santa Monica Mountains, coastal and chaparral regions of southern California. Santa Barbara, CA: Capra Press. In cooperation with: The California Native Plant Society. 239 p. 
51. Davis, Edwin A.; Kuester, James L.; Bagby, Marvin O. 1984. Biomass conversion to liquid fuels: potential of some Arizona chaparral brush and tree species. Nature. 307: 726-728. 
52. Davis, William B.; Taylor, Walter P. 1939. The bighorn sheep of Texas. Journal of Mammalogy. 20(4): 440-455. 
53. Dayton, William A. 1931. Important western browse plants. Misc. Publ. 101. Washington, DC: U.S. Department of Agriculture. 214 p. 
54. DeBano, Leonard F. 1990. Effects of fire on the soil resource in Arizona chaparral. In: Krammes, J. S., technical coordinator. Effects of fire management of southwestern natural resources: Proceedings of the symposium; 1988 November 15-17; Tucson, AZ. Gen. Tech. Rep. RM-191. Fort Collins, CO: U.S. Department of Agriculture, Forest Service, Rocky Mountain Forest and Range Experiment Station: 65-77. 
55. Denyes, H. Arliss. 1956. Natural terrestrial communities of Brewster County, Texas, with special reference to the distribution of the mammals. The American Midland Naturalist. 55(2): 289-320. 
56. Diamond, David D.; Riskind, David H.; Orzell, Steve L. 1987. A framework for plant community classification and conservation in Texas. Texas Journal of Science. 39(3): 203-221. 
57. Dick-Peddie, William A. 1993. New Mexico vegetation: Past, present, and future. Albuquerque, NM: University of New Mexico Press. 244 p. 
58. Dieterich, J. H.; Hibbert, A. R. 1990. Fire history in a small ponderosa pine stand surrounded by chaparral. In: Krammes, J. S., technical coordinator. Effects of fire management of southwestern natural resources: Proceedings of the symposium; 1988 November 15-17; Tucson, AZ. Gen. Tech. Rep. RM-191. Fort Collins, CO: U.S. Department of Agriculture, Forest Service, Rocky Mountain Forest and Range Experiment Station: 168-173. 
59. Dietz, Donald R.; Uresk, Daniel W.; Messner, Harold E.; McEwen, Lowell C. 1980. Establishment, survival, and growth of selected browse species in a ponderosa pine forest. Res. Pap. RM-219. Fort Collins, CO: U.S. Department of Agriculture, Forest Service, Rocky Mountain Forest and Range Experiment Station. 11 p. 
60. Dimeo, Art. 1977. Project record: An investigation of equipment for rejuvenating browse. ED&T 7080: Equipment to Rejuvenate Browse. Missoula, MT: U.S. Department of Agriculture, Forest Service, Equipment Development Center. 19 p. 
61. Dittberner, Phillip L.; Olson, Michael R. 1983. The plant information network (PIN) data base: Colorado, Montana, North Dakota, Utah, and Wyoming. FWS/OBS-83/86. Washington, DC: U.S. Department of the Interior, Fish and Wildlife Service. 786 p. 
62. Donnegan, Joseph A. 1999. Climatic and human influences on fire regimes in Pike National Forest. Boulder, CO: University of Colorado. 122 p. Dissertation. 
63. Donnegan, Joseph A.; Veblen, Thomas T.; Sibold, Jason S. 2001. Climatic and human influences on fire history in Pike National Forest, central Colorado. Canadian Journal of Forest Research. 31: 1526-1539. 
64. Dorn, Robert D. 1988. Vascular plants of Wyoming. Cheyenne, WY: Mountain West Publishing. 340 p. 
65. Duran, John C. 1973. Field investigations and energy determinations of stomach contents of Peromyscus boylii in the Granite Basin area, Yavapai County, Arizona. Greeley, CO: University of Northern Colorado. 62 p. Dissertation. 
66. Elmore, Francis H. 1944. Ethnobotany of the Navajo. Monograph Series: 1(7). Albuquerque, NM: University of New Mexico. 136 p. 
67. Erdman, James A. 1970. Pinyon-juniper succession after natural fires on residual soils of Mesa Verde, Colorado. Brigham Young University Science Bulletin: Biological Series. 11(2): 1-26. 
68. Erdman, James Allen. 1969. Pinyon-juniper succession after fires on residual soils of the Mesa Verde, Colorado. Boulder, CO: University of Colorado. 81 p. Dissertation. 
69. Eyre, F. H., ed. 1980. Forest cover types of the United States and Canada. Washington, DC: Society of American Foresters. 148 p. 
70. Ferguson, Robert B. 1983. Use of rosaceous shrubs for wildland plantings in the Intermountain West. In: Monsen, Stephen B.; Shaw, Nancy, comps. Managing Intermountain rangelands--improvement of range and wildlife habitats; Proceedings of symposia; 1981 September 15-17; Twin Falls, ID; 1982 June 22-24; Elko, NV. Gen. Tech. Rep. INT-157. Ogden, UT: U.S. Department of Agriculture, Forest Service, Intermountain Forest and Range Experiment Station: 136-149. 
71. Ferguson, Robert B.; Frischknecht, Neil C. 1985. Reclamation on Utah's Emery and Alton coal fields: techniques and plant materials. Res. Pap. INT-335. Ogden, UT: U.S. Department of Agriculture, Forest Service, Intermountain Forest and Range Experiment Station. 78 p. 
72. Floyd, M. Lisa; Romme, William H.; Hanna, David D. 2000. Fire history and vegetation pattern in Mesa Verde National Park, Colorado, USA. Ecological Applications. 10(6): 1666-1680. 
73. Foxx, Teralene S. 1996. Vegetation succession after the La Mesa Fire at Bandelier National Monument. In: Allen, Craig D., ed. Fire effects in southwestern forests: Proceedings, 2nd La Mesa fire symposium; 1994 March 29-31; Los Alamos, NM. RM-GTR-286. Fort Collins, CO: U.S. Department of Agriculture, Forest Service, Rocky Mountain Forest and Range Experiment Station: 47-69. 
74. Foxx, Teralene S.; Tierney, Gail D. 1987. Rooting patterns in the pinyon-juniper woodland. In: Everett, Richard L., compiler. Proceedings--pinyon-juniper conference; 1986 January 13-16; Reno, NV. Gen. Tech. Rep. INT-215. Ogden, UT: U.S. Department of Agriculture, Forest Service, Intermountain Research Station: 69-79. 
75. Franklin, Janet; Coulter, Charlotte L.; Rey, Sergio J. 2004. Change over 70 years in a southern California chaparral community related to fire history. Journal of Vegetation Science. 15(5): 701-710. 
76. Frost, William E. 1989. The Ellis Ranch Project: a case study in controlled burning. No. 891002. Fresno, CA: California Agricultural Technology Institute; San Joaquin Experimental Range. 11 p. 
77. Garrison, George A.; Bjugstad, Ardell J.; Duncan, Don A.; Lewis, Mont E.; Smith, Dixie R. 1977. Vegetation and environmental features of forest and range ecosystems. Agric. Handb. 475. Washington, DC: U.S. Department of Agriculture, Forest Service. 68 p. 
78. Gay, Charles W., Jr.; Dwyer, Don D. 1965. New Mexico range plants. Circular 374. Las Cruces, NM: New Mexico State University, Cooperative Extension Service. 85 p. 
79. Gese, Eric M.; Rongstad, Orrin J.; Mytton, William R. 1988. Home range and habitat use of coyotes in southeastern Colorado. Journal of Wildlife Management. 52(4): 640-646. 
80. Gibbens, R. P.; Schultz, A. M. 1962. Manipulation of shrub form and browse production in game range improvement. California Fish and Game. 48: 49-64. 
81. Gibbens, R. P.; Schultz, A. M. 1963. Brush manipulation on a deer winter range. California Fish and Game. 49(2): 95-118. 
82. Goodrich, Sherel. 1999. Multiple use management based on diversity of capabilities and values within pinyon-juniper woodlands. In: Monsen, Stephen B.; Stevens, Richard, compilers. Proceedings: ecology and management of pinyon-juniper communities within the Interior West: Sustaining and restoring a diverse ecosystem; 1997 September 15-18; Provo, UT. Proceedings RMRS-P-9. Ogden, UT: U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station: 164-171. 
83. Goodrich, Sherel; Neese, Elizabeth. 1986. Uinta Basin flora. Ogden, UT: U.S. Department of Agriculture, Forest Service, Intermountain Region, Ashley National Forest; U.S. Department of the Interior, Bureau of Land Management, Vernal District. 320 p. 
84. Goodwin, Gregory A. 1975. Seasonal food habits of mule deer in southeastern Wyoming. Research Note RM-287. Fort Collins, CO: U.S. Department of Agriculture, Forest Service, Rocky Mountain Forest and Range Experiment Station. 4 p. 
85. Gottfried, Gerald J.; Swetnam, Thomas W.; Allen, Craig D.; Betancourt, Julio L.; Chung-MacCoubrey, Alice L. 1995. Pinyon-juniper woodlands. In: Finch, Deborah M.; Tainter, Joseph A., eds. Ecology, diversity, and sustainability of the Middle Rio Grande Basin. Gen. Tech. Rep. RM-GTR-268. Fort Collins, CO: U.S. Department of Agriculture, Forest Service, Rocky Mountain Forest and Range Experiment Station: 95-132. 
86. Great Plains Flora Association. 1986. Flora of the Great Plains. Lawrence, KS: University Press of Kansas. 1392 p. 
87. Green, Lisle R. 1970. An experimental prescribed burn to reduce fuel hazard in chaparral. Res. Note PSW-216. Berkeley, CA: U.S. Department of Agriculture, Forest Service, Pacific Southwest Forest and Range Experiment Station. 6 p. 
88. Green, Lisle R. 1981. Burning by prescription in chaparral. Gen. Tech. Rep. PSW-51. Berkeley, CA: U.S. Department of Agriculture, Forest Service, Pacific Southwest Forest and Range Experiment Station. 36 p. 
89. Green, Lisle R.; Newell, Leonard A. 1982. Using goats to control brush regrowth on fuelbreaks. Gen. Tech. Rep. PSW-59. Berkeley, CA: U.S. Department of Agriculture, Forest Service, Pacific Southwest Forest and Range Experiment Station. 13 p. 
90. Greenwood, Larry R.; Brotherson, Jack D. 1978. Ecological relationships between pinyon-juniper and true mountain mahogany stands in the Uintah Basin, Utah. Journal of Range Management. 31(3): 164-167. 
91. Grissino-Mayer, Henri. 1995. Tree-ring reconstructions of climate and fire history at El Malpais National Monument, New Mexico. Tucson, AZ: University of Arizona. 407 p. Dissertation. 
92. Griswold, Sylvia M. 1936. Effect of alternate moistening and drying on germination of seeds of western range plants. Botanical Gazette. 98(2): 243-269. 
93. Gruell, G. E.; Loope, L. L. 1974. Relationships among aspen, fire, and ungulate browsing in Jackson Hole, Wyoming. Fort Collins, CO: U.S. Department of Agriculture, Forest Service, Rocky Mountain Forest and Range Experiment Station. 33 p. In cooperation with: U.S. Department of the Interior, National Park Service, Rocky Mountain Region. 
94. Guo, Qinfeng. 2001. Early post-fire succession in California chaparral: changes in diversity, density, cover and biomass. Ecological Research. 16: 471-485. 
95. Gutknecht, Kurt W. 1989. Xeriscaping: an alternative to thirsty landscapes. Utah Science. 50(4): 142-146. 
96. Haggerty, P. K. 1994. Damage and recovery in southern Sierra Nevada foothill oak woodland after a severe ground fire. Madrono. 41(3): 185-198. 
97. Halsey, Richard W. 2005. Chaparral, California's unknown wilderness. In: Halsey, Richard W. Fire, chaparral, and survival in southern California. San Diego, CA: Sunbelt Publications, Inc.: 1-30. 
98. Hamilton, John W.; Gilbert, Carl S. 1972. Composition of Wyoming range plants and soils. Research Journal No. 55. Laramie, WY: University of Wyoming, Agricultural Experiment Station. 14 p. 
99. Hanes, Ted L. 1971. Succession after fire in the chaparral of southern California. Ecological Monographs. 41(1): 27-52. 
100. Hanes, Ted L. 1981. California chaparral. In: Di Castri, F.; Goodall, D. W.; Specht, R. L., eds. Mediterranean-type shrublands. Amsterdam: Elsevier Science Publishers B.V: 139-174. 
101. Hanes, Ted L.; Friesen, Richard D.; Keane, Kathy. 1989. Alluvial scrub vegetation in coastal southern California. In: Protection, management, and restoration for the 1990's: Proceedings of the California Riparian Systems conference; 1988 September 22-24; Davis, CA. Gen. Tech. Rep. PSW-110. Berkeley, CA: U.S. Department of Agriculture, Forest Service, Pacific Southwest Forest and Range Experiment Station: 187-193. 
102. Hanes, Ted L.; Jones, Harold W. 1967. Postfire chaparral succession in southern California. Ecology. 48(2): 259-264. 
103. Hanks, Jess P.; Dick-Peddie, W. A. 1974. Vegetation patterns of the White Mountains, New Mexico. The Southwestern Naturalist. 18(4): 371-382. 
104. Harrington, H. D. 1964. Manual of the plants of Colorado. 2nd ed. Chicago: The Swallow Press, Inc. 666 p. 
105. Harrison, A. T.; Small, E.; Mooney, H. A. 1971. Drought relationships and distribution of two Mediterranean-climate California plant communities. Ecology. 52(5): 869-875. 
106. Haufler, Jonathan B.; Nagy, Julius G. 1984. Summer food habits of a small mammal community in the pinyon-juniper ecosystem. The Great Basin Naturalist. 44(1): 145-150. 
107. Hayes, Doris W.; Garrison, George A. 1960. Key to important woody plants of eastern Oregon and Washington. Agric. Handb. 148. Washington, DC: U.S. Department of Agriculture, Forest Service. 227 p. 
108. Hayward, C. Lynn. 1948. Biotic communities of the Wasatch chaparral, Utah. Ecological Monographs. 18: 473-506. 
109. Hellmers, H.; Horton, J. S.; Juhren, G.; O'Keefe, J. 1955. Root systems of some chaparral plants in southern California. Ecology. 36(4): 667-678. 
110. Hess, Karl; Alexander, Robert R. 1986. Forest vegetation of the Arapaho and Roosevelt National Forests in central Colorado: a habitat type classification. Res. Pap. RM-266. Fort Collins, CO: U.S. Department of Agriculture, Forest Service, Rocky Mountain Forest and Range Experiment Station. 48 p. 
111. Hess, Karl; Wasser, Clinton H. 1982. Grassland, shrubland, and forestland habitat types of the White River-Arapaho National Forest. Final Report. Fort Collins, CO: U.S. Department of Agriculture, Forest Service, Rocky Mountain Forest and Range Experiment Station. 335 p. 
112. Hickman, James C., ed. 1993. The Jepson manual: Higher plants of California. Berkeley, CA: University of California Press. 1400 p. 
113. Hill, Alison; Pieper, Rex D.; Southward, G. Morris. 1992. Habitat-type classification of the pinyon-juniper woodlands in western New Mexico. Bulletin 766. Las Cruces, NM: New Mexico State University, College of Agriculture and Home Economics, Agricultural Experiment Station. 80 p. 
114. Hitchcock, C. Leo; Cronquist, Arthur. 1961. Vascular plants of the Pacific Northwest. Part 3: Saxifragaceae to Ericaceae. Seattle, WA: University of Washington Press. 614 p. 
115. Hitchcock, C. Leo; Cronquist, Arthur. 1973. Flora of the Pacific Northwest. Seattle, WA: University of Washington Press. 730 p. 
116. Hoffman, George R.; Alexander, Robert R. 1987. Forest vegetation of the Black Hills National Forest of South Dakota and Wyoming: a habitat type classification. Res. Pap. RM-276. Fort Collins, CO: U.S. Department of Agriculture, Forest Service, Rocky Mountain Forest and Range Experiment Station. 48 p. 
117. Holland, Robert F. 1986. Preliminary descriptions of the terrestrial natural communities of California. Sacramento, CA: California Department of Fish and Game. 156 p. 
118. Horton, Howard, ed./comp. 1989. Interagency forage and conservation planting guide for Utah. Extension Circular 433. Logan, UT: Utah State University, Cooperative Extension Service. 67 p. 
119. Horton, Jerome S. 1960. Vegetation types of the San Bernardino Mountains. Tech. Rep. PSW-44. Berkeley, CA: U.S. Department of Agriculture, Forest Service, Pacific Southwest Forest and Range Experiment Station. 29 p. 
120. Horton, Jerome S.; Wright, John T. 1944. The wood rat as an ecological factor in southern California watersheds. Ecology. 25(3): 341-351. 
121. Howard, Gene S.; Rauzi, Frank; Schuman, Gerald E. 1979. Woody plant trials at six mine reclamation sites in Wyoming and Colorado. Production Res. Rep. PRR 177/1/79. Washington, DC: U.S. Department of Agriculture. 14 p. 
122. Huber, Allen; Goodrich, Sherel; Anderson, Kim. 1999. Diversity with successional status in the pinyon-juniper/mountain mahogany/bluebunch wheatgrass community type near Dutch John, Utah. In: Monsen, Stephen B.; Stevens, Richard, compilers. Proceedings: ecology and management of pinyon-juniper communities within the Interior West: Sustaining and restoring a diverse ecosystem; 1997 September 15-18; Provo, UT. Proceedings RMRS-P-9. Ogden, UT: U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station: 114-117. 
123. Humphrey, Robert R. 1960. Forage production on Arizona ranges. V. Pima, Pinal and Santa Cruz Counties. Bulletin 502. Tucson, AZ: University of Arizona, Agricultural Experiment Station. 137 p. 
124. Idaho Native Plant Society. 2004. The Idaho Native Plant Society rare plant list: State rare species list, [Online]. In: Results of the 20th annual Idaho rare plant conference. Idaho Native Plant Society (Producer). Available: http://www.idahonativeplants.org/rpc/RarePlantList.aspx [2005, January 24]. 
125. Institute for Land Rehabilitation. 1979. Selection, propagation, and field establishment of native plant species on disturbed arid lands. Bulletin 500. Logan, UT: Utah State University, Agricultural Experiment Station. 49 p. 
126. Johnson, E. W. 1963. Ornamental shrubs for the southern Great Plains. Farmer's Bull. 2025. Washington, DC: U.S. Department of Agriculture. 62 p. 
127. Jorgensen, Kent R.; Stevens, Richard. 2004. Seed collection, cleaning, and storage. In: Monsen, Stephen B.; Stevens, Richard; Shaw, Nancy L., comps. Restoring western ranges and wildlands. Gen. Tech. Rep. RMRS-GTR-136-vol-3. Fort Collins, CO: U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station: 699-716. 
128. Judd, B. Ira. 1962. Principal forage plants of southwestern ranges. Stn. Pap. No. 69. Fort Collins, CO: U.S. Department of Agriculture, Forest Service, Rocky Mountain Forest and Range Experiment Station. 93 p. 
129. Juhren, Gustaf; Pole, Rupert; O'Keefe, James. 1955. Conversion of brush to grass on a burned chaparral area. Journal of Forestry. 53(5): 348-351. 
130. Julander, Odell. 1937. Utilization of browse by wildlife. Transactions, 2nd North American Wildlife Conference: 276-287. 
131. Julander, Odell; Robinette, W. Leslie. 1950. Deer and cattle range relationships on Oak Creek range in Utah. Journal of Forestry. 48(6): 410-415. 
132. Kartesz, John T.; Meacham, Christopher A. 1999. Synthesis of the North American flora (Windows Version 1.0), [CD-ROM]. Available: North Carolina Botanical Garden. In cooperation with: The Nature Conservancy, Natural Resources Conservation Service, and U.S. Fish and Wildlife Service [2001, January 16]. 
133. Kartesz, John Thomas. 1988. A flora of Nevada. Reno, NV: University of Nevada. 1729 p. [In 2 volumes]. Dissertation. 
134. Kearney, Thomas H.; Peebles, Robert H.; Howell, John Thomas; McClintock, Elizabeth. 1960. Arizona flora. 2nd ed. Berkeley, CA: University of California Press. 1085 p. 
135. Keeley, J. E.; Brooks, A.; Bird, T.; Cory, S.; Parker, H.; Usinger, E. 1986. Demographic structure of chaparral under extended fire-free conditions. In: DeVries, Johannes J., ed. Proceedings of the chaparral ecosystems research conference; 1985 May 16-17; Santa Barbara, CA. Report No. 2. Davis, CA: University of California, California Water Resources Center: 133-137. 
136. Keeley, Jon E. 1981. Reproductive cycles and fire regimes. In: Mooney, H. A.; Bonnicksen, T. M.; Christensen, N. L.; Lotan, J. E.; Reiners, W. A., tech. coords. Fire regimes and ecosystem properties: Proceedings of the conference; 1978 December 11-15; Honolulu, HI. Gen. Tech. Rep. WO-26. Washington, DC: U.S. Department of Agriculture, Forest Service: 231-277. 
137. Keeley, Jon E. 1986. Resilience of Mediterranean shrub communities to fires. In: Dell, B.; Hopkins, A. J. N.; Lamont B. B., eds. Resilience in Mediterranean-type ecosystems. Dordrecht, The Netherlands: Dr. W. Junk Publishers: 95-112. 
138. Keeley, Jon E. 1987. Role of fire in seed germination of woody taxa in California chaparral. Ecology. 68(2): 434-443. 
139. Keeley, Jon E. 1991. Fire management for maximum biodiversity of California chaparral. In: Nodvin, Stephen C.; Waldrop, Thomas A., eds. Fire and the environment: ecological and cultural perspectives: Proceedings of an international symposium; 1990 March 20-24; Knoxville, TN. Gen. Tech. Rep. SE-69. Asheville, NC: U.S. Department of Agriculture, Forest Service, Southeastern Forest Experiment Station: 11-14. 
140. Keeley, Jon E. 1992. Demographic structure of California chaparral in the long-term absence of fire. Vegetation Science. 3(1): 79-90. 
141. Keeley, Jon E. 1992. Recruitment of seedlings and vegetative sprouts in unburned chaparral. Ecology. 73(4): 1194-1208. 
142. Keeley, Jon E.; Fotheringham, C. J. 2003. Impact of past, present, and future fire regimes on North American Mediterranean shrublands. In: Veblen, Thomas T.; Baker, William L.; Montenegro, Gloria; Swetnam, Thomas W., eds. Fire and climatic change in temperate ecosystems of the western Americas. Ecological Studies, Vol. 160. New York: Springer: 218-262. 
143. Keeley, Jon E.; Fotheringham, C. J.; Baer-Keeley, Melanie. 2006. Demographic patterns of postfire regeneration in Mediterranean-climate shrublands of California. Ecological Monographs. 76(2): 235-255. 
144. Keeley, Jon E.; Keeley, Sterling C. 1981. Post-fire regeneration of southern California chaparral. American Journal of Botany. 68(4): 524-530. 
145. Keeley, Jon E.; Keeley, Sterling C. 1986. Chaparral and wildfires. Fremontia. 14(3): 18-21. 
146. Keeley, Jon E.; Keeley, Sterling C. 1988. Chaparral. In: Barbour, Michael G.; Billings, William Dwight, eds. North American terrestrial vegetation. Cambridge; New York: Cambridge University Press: 165-207. 
147. Keeley, Jon E.; Pfaff, Anne H.; Safford, Hugh D. 2005. Fire suppression impacts on postfire recovery of Sierra Nevada chaparral shrublands. International Journal of Wildland Fire. 14: 255-265. 
148. Keeley, Jon E.; Zedler, Paul H.; Zammit, Charles A.; Stohlgren, Thomas J. 1989. Fire and demography. In: Keeley, Sterling C., ed. The California chaparral: Paradigms reexamined. Science Series No. 34. Los Angeles, CA: Natural History Museum of Los Angeles County: 151-153. 
149. Kennedy, Kathryn L. 1983. A habitat type classification of the pinyon-juniper woodlands of the Lincoln National Forest, New Mexico. In: Moir, W. H.; Hendzel, Leonard, tech. coords. Proceedings of the workshop on southwestern habitat types; 1983 April 6-8; Albuquerque, NM. Albuquerque, NM: U.S. Department of Agriculture, Forest Service, Southwestern Region: 54-61. 
150. Kinucan, Edith Seyfert. 1965. Deer utilization of postfire chaparral shrubs and fire history of the San Gabiel Mountains. Los Angeles, CA: California State College, Los Angeles. 61 p. Thesis. 
151. Kitchen, Stanley G. 2004. Cercocarpus montanus. In: Francis, John K., ed. Wildland shrubs of the United States and its territories: thamnic descriptions: volume 1. Gen. Tech. Rep. IITF-GTR-26. San Juan, PR: U.S. Department of Agriculture, Forest Service, International Institute of Tropical Forestry; Fort Collins, CO: U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station: 178-180. 
152. Kitchen, Stanley G.; Meyer, Susan E. 1990. Seed dormancy in two species of mountain mahogany (Cercocarpus ledifolius and Cercocarpus montanus). In: Johnson, Kendall L., ed. Proceedings, 5th Utah shrub ecology workshop: The genus Cercocarpus; 1988 July 13-14; Logan, UT. Logan, UT: Utah State University, College of Natural Resources: 27-41. 
153. Kitchen, Stanley G. [In press]. Cercocarpus Kunth--mountain-mahogany, [Online]. In: Bonner, Franklin T.; Nisley, Rebecca G.; Karrfait, R. P.; coords. Woody plant seed manual. Agric. Handbook 727. Washington, DC: U.S. Department of Agriculture, Forest Service (Producer). Available: http://www.nsl.fs.fed.us/wpsm/Cercocarpus.pdf [2006, August 18]. 
154. Kittams, Walter H. 1973. Effect of fire on vegetation of the Chihuahuan Desert region. In: Proceedings, annual Tall Timbers fire ecology conference; 1972 June 8-9; Lubbock, Texas. No. 12. Tallahassee, FL: Tall Timbers Research Station: 427-444. 
155. Kittams, Walter H.; Evans, Stanley L.; Cooke, Derrick C. 1979. Food habits of mule deer on foothills of Carlsbad Caverns National Park. In: Genoways, Hugh H.; Baker, Robert J., eds. Biological investigations in the Guadalupe Mountains National Park: Proceedings of a symposium; 1975 April 4-5; Lubbock, TX. Proceedings and Transactions Series No. 4. Washington, DC: U.S. Department of the Interior, National Park Service: 403-426. 
156. Kronemann, Loren A.; Britton, Carlton. 1982. Nutritional changes of selected wildlife browse after fire. In: Research highlights--1982: Noxious brush and weed control; range and wildlife management. Volume 13. Lubbock, TX: Texas Tech University, Department of Range and Wildlife Management: 70-71. 
157. Kruckeberg, A. R. 1982. Gardening with native plants of the Pacific Northwest. Seattle: University of Washington Press. 252 p. 
158. Krysl, Leslie J.; Moody, John D.; Simpson, C. David. 1979. Mule deer food habits and preferences in Guadalupe Mountains National Park, Texas. In: Sosebee, Ronald E.; Wright, Henry A., eds. Research highlights--1979: Noxious brush and weed control; range and wildlife management. Volume 10. Lubbock, TX: Texas Tech University, College of Agricultural Sciences: 48-49. 
159. Krysl, Leslie J.; Simpson, C. David; Gray, Gary G. 1979. Dietary overlap of sympatric barbary sheep and mule deer in Palo Duri Canyon, Texas. In: Sosebee, Ronald E.; Wright, Henry A.; eds. Research highlights--1979: Noxious brush and weed control; range and wildlife management. Volume 10. Lubbock, TX: Texas Tech University, College of Agricultural Sciences: 51-52. 
160. Kuchler, A. W. 1964. Manual to accompany the map of potential vegetation of the conterminous United States. Special Publication No. 36. New York: American Geographical Society. 77 p. 
161. Kufeld, Roland C.; Bowden, David C.; Schrupp, Donald L. 1988. Habitat selection and activity patterns of female mule deer in the Front Range, Colorado. Journal of Range Management. 41(6): 515-522. 
162. Kufeld, Roland C.; Stevens, Marilyn L.; Bowden, David C. 1985. Site variation in forage qualities of mountain mahogany and serviceberry. Journal of Range Management. 38(5): 458-460; 1985. 
163. Kufeld, Roland C.; Wallmo, O. C.; Feddema, Charles. 1973. Foods of the Rocky Mountain mule deer. Res. Pap. RM-111. Fort Collins, CO: U.S. Department of Agriculture, Forest Service, Rocky Mountain Forest and Range Experiment Station. 31 p. 
164. Kummerow, J.; Mangan, R. 1981. Root systems in Quercus dumosa Nutt. dominated chaparral in southern California. Acta Oecologica. 2(16): 177-188. 
165. Landis, Thomas D.; Simonich, Edward J. 1984. Producing native plants as container seedlings. In: Murphy, Patrick M., comp. The challenge of producing native plants for the Intermountain area: Proceedings: Intermountain Nurseryman's Association 1983 conference; 1983 August 8-11; Las Vegas, NV. Gen. Tech. Rep. INT-168. Ogden, UT: U.S. Department of Agriculture, Forest Service, Intermountain Forest and Range Experiment Station: 16-25. 
166. Larson, Milo; Moir, W. H. 1987. Forest and woodland habitat types (plant associations) of northern New Mexico and northern Arizona. 2nd ed. Albuquerque, NM: U.S. Department of Agriculture, Forest Service, Southwestern Region. 90 p. 
167. Lathrop, Earl W.; Martin, Bradford D. 1982. Response of understory vegetation to prescribed burning in yellow pine forests of Cuyamaca Rancho State Park, California. Aliso. 10(2): 329-343. 
168. Laven, R. D.; Omi, P. N.; Wyant, J. G.; Pinkerton, A. S. 1980. Interpretation of fire scar data from a ponderosa pine ecosystem in the central Rocky Mountains, Colorado. In: Stokes, Marvin A.; Dieterich, John H., tech. coords. Proceedings of the fire history workshop; 1980 October 20-24; Tucson, AZ. Gen. Tech. Rep. RM-81. Fort Collins, CO: U.S. Department of Agriculture, Forest Service, Rocky Mountain Forest and Range Experiment Station: 46-49. 
169. Leach, Howard R.; Hiehle, Jack L. 1956. Food habits of the Tehama deer herd. California Fish and Game. 43: 161-178. 
170. Lewis, Henry T. 1973. Patterns of Indian burning in California: Ecology and ethnohistory. Ballena Press Anthropological Papers No. 1. Ramona, CA: Ballena Press. 101 p. 
171. Liang, Li-Ming; Omi, P. N. 2003. True mountain mahogany sprouting behavior following fire, [Online]. In: Proceedings, 2nd international wildland fire ecology and fire management congress; 5th symposium on fire and forest meteorology; 2003 November 16-20; Orlando, FL. Boston, MA: American Meteorology Society (Producer). Available: http://ams.confex.com/ams/FIRE2003/techprogram/paper_65787.htm [2006, October 27]. 
172. Little, Elbert L., Jr. 1950. Southwestern trees: A guide to the native species of New Mexico and Arizona. Agriculture Handbook No. 9. Washington, DC: U.S. Department of Agriculture, Forest Service. 109 p. 
173. Loik, Michael E.; St. Onge, Christine D.; Rogers, Jane. 2000. Post-fire recruitment of Yucca brevifolia and Yucca schidigera in Joshua Tree National Park, California. In: Keeley, Jon E.; Baer-Keeley, Melanie; Fotheringham, C. J., eds. 2nd interface between ecology and land development in California. U.S. Geological Survey: Open-File Report 00-62. Sacramento, CA: U.S. Department of the Interior, Geological Survey, Western Ecological Research Center: 79-85. 
174. Mahgoub, El Fatih; Pieper, Rex D.; Holechek, Jerry L.; Wright, Jimmie D.; Howard, V. W., Jr. 1987. Botanical content of mule deer diets in south-central New Mexico. New Mexico Journal of Science. 27(1): 21-27. 
175. Mahgoub, El Fatih; Pieper, Rex D.; Ortiz, Melchor. 1988. Use of leader lengths and diameters to estimate production and utilization of Cercocarpus breviflorus. Journal of Range Management. 41(2): 153-155. 
176. Malanson, George P.; O'Leary, John F. 1985. Effects of fire and habitat on post-fire regeneration in Mediterranean-type ecosystems: Ceanothus spinosus chaparral and Californian coastal sage scrub. Acta Oecologica. 6(20): 169-181. 
177. Martin, Floyd L. 1950. A revision of Cercocarpus. Brittonia. 7(2): 91-111. 
178. Martin, William C.; Hutchins, Charles R. 1981. A flora of New Mexico. Volume 2. Germany: J. Cramer. 2589 p. 
179. McArthur, E. Durant; Giunta, Bruce C.; Plummer, A. Perry. 1977. Shrubs for restoration of depleted range and disturbed areas. Utah Science. 35: 28-33. 
180. McCandliss, David S. 2002. Prescribed burning in the Kings River Ecosystems Project area: lessons learned. In: Verner, Jared, tech. ed. Proceedings of a symposium on the Kings River Sustainable Forest Ecosystems Project: progress and current status; 1998 January 26; Clovis, CA. Gen. Tech. Rep. PSW-GTR-183. Albany, CA: U.S. Department of Agriculture, Forest Service, Pacific Southwest Research Station: 37-45. 
181. McCulloch, Clay Y. 1973. Part I: Seasonal diets of mule and white-tailed deer. In: Deer nutrition in Arizona chaparral and desert habitats. Special Report No. 3: Federal Aid in Wildlife Restoration Act Project W-78-R. Phoenix, AZ: Arizona Game and Fish Department, Research Division: 1-37. In cooperation with: U.S. Department of Agriculture, Forest Service, Rocky Mountain Forest and Range Experiment Station. 
182. McIntosh, Arthur C. 1931. A botanical survey of the Black Hills of South Dakota. Black Hills Engineer. 19(3): 159-276. 
183. Meinecke, E. P. 1929. Quaking aspen: A study in applied forest pathology. Tech. Bull. No. 155. Washington, DC: U.S. Department of Agriculture. 34 p. 
184. Minnich, Richard A. 1976. Vegetation of the San Bernardino Mountains. In: Latting, June, ed. Symposium proceedings: plant communities of southern California; 1974 May 4; Fullerton, CA. Special Publication No. 2. Berkeley, CA: California Native Plant Society: 99-124. 
185. Minnich, Richard A. 1987. Fire behavior in southern California chaparral before fire control: the Mount Wilson burns at the turn of the century. Annals of the Association of American Geographers. 77(4): 599-618. 
186. Minnich, Richard A. 1999. Vegetation, fire regimes, and forest dynamics. In: Miller, P. R.; McBride, J. R., eds. Oxidant air pollution impacts in the montane forests of southern California: a case study of the San Bernardino Mountains. Ecological Studies: Analysis and Synthesis, Vol. 134. New York: Springer-Verlag: 44-80. 
187. Minnich, Richard A.; Bahre, Conrad J. 1995. Wildland fire and chaparral succession along the California-Baja California boundary. International Journal of Wildland Fire. 5(1): 13-24. 
188. Minnich, Richard A.; Franco-Vizcaino, Ernesto. 1997. Mediterranean vegetation of northern Baja California. Fremontia. 25(3): 3-12. 
189. Mirov, N. T. 1936. Germination behavior of some California plants. Ecology. 17(4): 667-672. 
190. Mitchell, Jerry M. 1984. Fire management action plan: Zion National Park, Utah. Record of Decision. 73 p. Salt Lake City, UT: U.S. Department of the Interior, National Park Service. On file with: U.S. Department of Agriculture, Forest Service, Intermountain Research Station, Fire Sciences Laboratory, Missoula, MT. 
191. Moir, W. H.; Carleton, J. O. 1987. Classification of pinyon-juniper (p-j) sites on national forests in the Southwest. In: Everett, Richard L., compiler. Proceedings--pinyon-juniper conference; 1986 January 13-16; Reno, NV. Gen. Tech. Rep. INT-215. Ogden, UT: U.S. Department of Agriculture, Forest Service, Intermountain Research Station: 216-226. 
192. Moir, William H. 1982. A fire history of the High Chisos, Big Bend National Park, Texas. The Southwestern Naturalist. 27(1): 87-98. 
193. Monsen, Stephen B. 1987. Shrub selections for pinyon-juniper plantings. In: Everett, Richard L., compiler. Proceedings--pinyon-juniper conference; 1986 January 13-16; Reno, NV. Gen. Tech. Rep. INT-215. Ogden, UT: U.S. Department of Agriculture, Forest Service, Intermountain Research Station: 316-329. 
194. Monsen, Stephen B.; McArthur, E. Durant. 1985. Factors influencing establishment of seeded broadleaf herbs and shrubs following fire. In: Sanders, Ken; Durham, Jack, eds. Rangeland fire effects: A symposium: Proceedings; 1984 November 27-29; Boise, ID. Boise, ID: U.S. Department of the Interior, Bureau of Land Management, Idaho State Office: 112-124. 
195. Monsen, Stephen B.; Stevens, Richard. 2004. Seedbed preparation and seeding practices. In: Monsen, Stephen B.; Stevens, Richard; Shaw, Nancy L., comps. Restoring western ranges and wildlands. Gen. Tech. Rep. RMRS-GTR-136-vol-1. Fort Collins, CO: U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station: 121-154. 
196. Montgomery, Kenneth Reid. 1976. Ether extractives and flammability of Mediterranean-type shrubs. Pomona, CA: California Polytechnic University, Pomona. 38 p. Thesis. 
197. Morrison, Peter H.; Swanson, Frederick J. 1990. Fire history and pattern in a Cascade Range landscape. Gen. Tech. Rep. PNW-GTR-254. Portland, OR: U.S. Department of Agriculture, Forest Service, Pacific Northwest Research Station. 77 p. 
198. Mouat, David A. 1974. Relationships between vegetation and terrain variables in southeastern Arizona. Corvallis, OR: Oregon State University. 242 p. Thesis. 
199. Mower, Kerry J.; Smith, H. Duane. 1989. Diet similarity between elk and deer in Utah. The Great Basin Naturalist. 49(4): 552-555. 
200. Munz, Philip A.; Keck, David D. 1973. A California flora and supplement. Berkeley, CA: University of California Press. 1905 p. 
201. Neff, Don J. 1974. Forage preferences of trained deer on the Beaver Creek watersheds. Special Report No. 4. Phoenix, AZ: Arizona Game and Fish Department. 61 p. 
202. Nelle, Stephan A. 1996. Management and use of browse in the Edwards Plateau of Texas. In: Barrow, Jerry R.; McArthur, E. Durant; Sosebee, Ronald E.; Tausch, Robin J., compilers. Proceedings: shrubland ecosystem dynamics in a changing environment; 1995 May 23-25; Las Cruces, NM. Gen. Tech. Rep. INT-GTR-338. Ogden, UT: U.S. Department of Agriculture, Forest Service, Intermountain Research Station: 151-155. 
203. Nelle, Stephan A. 2001. Ecological implications of using goats for control of juniper in Texas. In: McArthur, E. Durant; Fairbanks, Daniel J., compilers. Shrubland ecosystem genetics and biodiversity: proceedings; 2000 June 13-15; Provo, UT. Proc. RMRS-P-21. Ogden, UT: U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station: 352-355. 
204. Nevada Natural Heritage Program. 2003. National vegetation classification for Nevada [NVC], [Online]. Carson City, NV: Nevada Department of Conservation and Natural Resources (Producer). Available: http://heritage.nv.gov/ecology/nv_nvc.htm [2005, November 3]. 
205. Nixon, E. S. 1977. A mountain cercocarpus population--revisited. The Great Basin Naturalist. 37(1): 97-99. 
206. Northcutt, Bennett Earl. 1978. The plant ecology of Butler Wash, southeastern Utah. Boulder, CO: University of Colorado. 135 p. Thesis. 
207. Northington, David K.; Burgess, Tony L. 1979. Summary of the vegetative zones of the Guadalupe Mountains National Park, Texas. In: Genoways, Hugh H.; Baker, Robert J., eds. Biological investigations in the Guadalupe Mountains National Park: Proceedings of a symposium; 1975 April 4-5; Lubbock, TX. Proceedings and Transactions Series No. 4. Washington, DC: U.S. Department of the Interior, National Park Service: 51-57. 
208. O'Leary, John Francis. 1984. Environmental factors influencing postburn vegetation in a southern California shrubland. Los Angeles, CA: University of California. 92 p. Dissertation. 
209. Olson, Rich. 1992. Mule deer habitat requirements and management in Wyoming. B-965. Laramie, WY: University of Wyoming, Cooperative Extension Service. 15 p. 
210. Olson, Rich. 1992. White-tailed deer habitat requirements and management in Wyoming. B-964. Laramie, WY: University of Wyoming, Cooperative Extension Service. 17 p. 
211. Olson, Thomas E.; Knopf, Fritz L. 1988. Patterns of relative diversity within riparian small mammal communities, Platte River watershed, Colorado. In: Szaro, Robert C.; Severson, Kieth E.; Patton, David R., technical coordinators. Management of amphibians, reptiles, and small mammals in North America: Proceedings of the symposium; 1988 July 19-21; Flagstaff, AZ. Gen. Tech. Rep. RM-166. Fort Collins, CO: U.S. Department of Agriculture, Forest Service, Rocky Mountain Forest and Range Experiment Station: 379-386. 
212. Ott, Jeffrey E.; McArthur, E. Durant; Sanderson, Stewart C. 2001. Plant community dynamics of burned and unburned sagebrush and pinyon-juniper vegetation in west-central Utah. In: McArthur, E. Durant; Fairbanks, Daniel J., compilers. Shrubland ecosystem genetics and biodiversity: proceedings; 2000 June 13-15; Provo, UT. Proc. RMRS-P-21. Ogden, UT: U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station: 177-191. 
213. Overby, S. T.; Perry, H. M. 1996. Direct effects of prescribed fire on available nitrogen and phosphorus in an Arizona chaparral watershed. Arid Soil and Research Rehabilitation. 10(4): 347-357. 
214. Parker, Karl G. 1975. Some important Utah range plants. Extension Service Bulletin EC-383. Logan, UT: Utah State University. 174 p. 
215. Parsons, David J. 1981. The historical role of fire in the foothill communities of Sequoia National Park. Madrono. 28(3): 111-120. 
216. Payne, Gene F. 1973. Vegetative rangeland types in Montana. Bull. 671. Bozeman, MT: Montana State University, Montana Agricultural Experiment Station. 15 p. 
217. Paysen, Timothy E.; Ansley, R. James; Brown, James K.; Gottfried, Gerald J.; Haase, Sally M.; Harrington, Michael G.; Narog, Marcia G.; Sackett, Stephen S.; Wilson, Ruth C. 2000. Fire in western shrubland, woodland, and grassland ecosystems. In: Brown, James K.; Smith, Jane Kapler, eds. Wildland fire in ecosystems: Effects of fire on flora. Gen. Tech. Rep. RMRS-GTR-42-volume 2. Ogden, UT: U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station: 121-159. 
218. Philbrick, Ralph N., Haller, J. R. 1977. The southern California islands. In: Barbour, Michael G.; Malor, Jack, eds. Terrestrial vegetation of California. New York: John Wiley and Sons: 893-906. 
219. Piatt, J. R. 1973. Seed size affects germination of true mountainmahogany. Journal of Range Management. 26(3): 231-232. 
220. Plumb, T. R. 1961. Sprouting of chaparral by December after a wildfire in July. Technical Paper 57. Berkeley, CA: U.S. Department of Agriculture, Forest Service, Pacific Southwest Forest and Range Experiment Station. 12 p. 
221. Plummer, A. Perry. 1977. Revegetation of disturbed Intermountain area sites. In: Thames, J. C., ed. Reclamation and use of disturbed lands of the Southwest. Tucson, AZ: University of Arizona Press: 302-337. 
222. Plummer, A. Perry; Christensen, Donald R.; Monsen, Stephen B. 1968. Restoring big-game range in Utah. Publ. No. 68-3. Ephraim, UT: Utah Division of Fish and Game. 183 p. 
223. Powell, A. Michael. 1988. Trees and shrubs of Trans-Pecos Texas: Including Big Bend and Guadalupe Mountains National Parks. Big Bend National Park, TX: Big Bend Natural History Association. 536 p. 
224. Prenzlow, E. J.; Gilbert, D. L.; Glover, F. A. 1968. Some behavior patterns of the pronghorn. Special Report No. 17/GFP-R-S 17. Denver, CO: Colorado Department of Game, Fish and Parks. 16 p. 
225. Rainier Seeds, Inc. 2003. Catalog, [Online]. Davenport, WA: Rainer Seeds, Inc., (Producer). Available: http://www.rainerseeds.com [2003, February 14]. 
226. Raunkiaer, C. 1934. The life forms of plants and statistical plant geography. Oxford: Clarendon Press. 632 p. 
227. Ream, Robert Ray. 1964. The vegetation of the Wasatch Mountains, Utah and Idaho. Madison, WI: University of Wisconsin. 178 p. Dissertation. 
228. Reeves, Timothy. 1976. Vegetation and flora of Chiricahua National Monument, Cochise County, Arizona. Tempe, AZ: Arizona State University. 180 p. Thesis. 
229. Reynolds, Hudson G. 1962. Some characteristics and uses of Arizona's major plant communities. Journal of the Arizona Academy of Science. 2: 62-71. 
230. Reynolds, Hudson G. 1964. Elk and deer habitat use of a pinyon-juniper woodland in southern New Mexico. In: Trefethen, James B., ed. Transactions, 29th North American wildlife and natural resources conference; 1964 March 9-11; Las Vegas, NV. Washington, DC: Wildlife Management Institute: 438-444. 
231. Reynolds, Hudson G. 1967. Chemical constituents and deer use of some crown sprouts in Arizona chaparral. Journal of Forestry. 65(12): 905-908. 
232. Reynolds, Hudson G.; Johnson, R. Roy. 1964. Habitat relations of vertebrates of the Sierra Ancha Experimental Forest. Res. Pap. RM-4. Fort Collins, CO: U.S. Department of Agriculture, Forest Service, Rocky Mountain Forest and Range Experiment Station. 16 p. 
233. Richter, Rebecca; Stromberg, Juliet C. 2005. Soil seed banks of two montane riparian areas: implications for restoration. Biodiversity and Conservation. 14(4): 993-1016. 
234. Riegel, Gregg M.; Smith, Bradley G.; Franklin, Jerry F. 1992. Foothill oak woodlands of the interior valleys of southwestern Oregon. Northwest Science. 66(2): 66-76. 
235. Rieseberg, Loren H. 1988. Saving California's rarest tree--Catalina mahogany. Plant Conservation. 3(1): 8. 
236. Riggan, Philip J.; Dunn, Paul H. 1982. Harvesting chaparral biomass for energy--an environmental assessment. In: Conrad, C. Eugene; Oechel, Walter C., technical coordinators. Proceedings of the symposium on dynamics and management of Mediterranean-type ecosystems; 1981 June 22-26; San Diego, CA. Gen. Tech. Rep. PSW-58. Berkeley, CA: U.S. Department of Agriculture, Forest Service, Pacific Southwest Forest and Range Experiment Station: 149-157. 
237. Riordan, Laurence E. 1958. Differences in range vegetation resulting from grazing by deer, cattle, and sheep. In: Proceedings: Society of American Foresters meeting; 1957 November 10-13; Syracuse, NY. Washington, DC: Society of American Foresters: 147-151. 
238. Ripple, William J. 1994. Historic spatial patterns of old forests in western Oregon. Journal of Forestry. 92(11): 45-49. 
239. Risenhoover, Kenneth L.; Bailey, James A. 1985. Foraging ecology of mountain sheep: implications for habitat management. Journal of Range Management. 49(3): 797-804. 
240. Rominger, Eric M.; Dale, Alan R.; Bailey, James A. 1988. Shrubs in the summer diet of Rocky Mountain bighorn sheep. Journal of Wildlife Management. 52(1): 47-50. 
241. Romme, William H.; Jamieson, David W.; Redders, Jeffery S.; Bigsby, Ginger; Lindsey, J. Page; Kendall, Deborah; Cowen, Robert; Kreykes, Thomas; Spencer, Albert W.; Ortega, Joseph C. 1992. Old-growth forests of the San Juan National Forest in southwestern Colorado. In: Kaufmann, Merrill R.; Moir, W. H.; Bassett, Richard L., tech. coords. Old-growth forests in the Southwest and Rocky Mountain regions: Proceedings of a workshop; 1992 March 9-13; Portal, AZ. Gen. Tech. Rep. RM-213. Fort Collins, CO: U.S. Department of Agriculture, Forest Service, Rocky Mountain Forest and Range Experiment Station: 154-165. 
242. Roper, Donald L. 1991. Mineral nutrient dynamics of four native shrub species in central Utah. Salt Lake City, UT: Brigham Young University. 94 p. Dissertation. 
243. Rosner, Lee S.; Harrington, John T.; Dreesen, David R.; Murray, Leigh. 2003. Overcoming dormancy in New Mexico mountain mahogany seed collections. Journal of Range Management. 56(2): 198-202. 
244. Rowland, M. M.; Alldredge, A. W.; Ellis, J. E.; Weber, B. J.; White, G. C. 1983. Comparative winter diets of elk in New Mexico. Journal of Wildlife Management. 47(4): 924-932. 
245. Sampson, Arthur W.; Jespersen, Beryl S. 1963. California range brushlands and browse plants. Berkeley, CA: University of California, Division of Agricultural Sciences; California Agricultural Experiment Station, Extension Service. 162 p. 
246. Sandoval, Leonard; Holechek, Jerry; Biggs, James; Valdez, Raul; VanLeeuwen, Dawn. 2005. Elk and mule deer diets in north-central New Mexico. Rangeland Ecology & Management. 58(4): 366-372. 
247. Schneider, Rick E.; Faber-Langendoen, Don; Crawford, Rex C.; Weakley, Alan S. 1997. The status of biodiversity in the Great Plains: Great Plains vegetation classification. Supplemental Document 1. In: Ostlie, Wayne R.; Schneider, Rick E.; Aldrich, Janette Marie; Faust, Thomas M.; McKim, Robert L. B.; Chaplin, Stephen J., compilers. The status of biodiversity in the Great Plains, [Online]. Arlington, VA: The Nature Conservancy (Producer). 75 p. Available: http://conserveonline.org/docs/2005/02/greatplains_vegclass_97.pdf [2006 May 16]. On file with: U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory, Missoula, MT. 
248. Schultz, Brad W. 1987. Ecology of curlleaf mountain mahogany (Cercocarpus ledifolius) in western and central Nevada: population structure and dynamics. Reno, NV: University of Nevada. 111 p. Thesis. 
249. Seklecki, Mariette T.; Grissino-Mayer, Henri D.; Swetnam, Thomas W. 1996. Fire history and the possible role of Apache-set fires in the Chiricahua Mountains of southeastern Arizona. In: Ffolliott, Peter F.; DeBano, Leonard F.; Baker, Malchus B., Jr.; Gottfried, Gerald J.; Solis-Garza, Gilberto; Edminster, Carleton B.; Neary, Daniel G.; Allen, Larry S.; Hamre, R. H., tech. coords. Effects of fire on Madrean Province ecosystems: a symposium proceedings; 1996 March 11-15; Tucson, AZ. Gen. Tech. Rep. RM-GTR-289. Fort Collins, CO: U.S. Department of Agriculture, Forest Service, Rocky Mountain Forest and Range Experiment Station: 238-246. 
250. Shaw, Nancy L.; Monsen, Stephen B.; Stevens, Richard. 2004. Rosaceous shrubs. In: Monsen, Stephen B.; Stevens, Richard; Shaw, Nancy L. Restoring western ranges and wildlands. Gen. Tech. Rep. RMRS-GTR-136-vol-2. Fort Collins, CO: U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station: 539-596. 
251. Shepherd, Harold R. 1971. Effects of clipping on key browse species in southwestern Colorado. Technical Publication Number 28. [Denver, CO]: Colorado Division of Game, Fish and Parks. 104 p. 
252. Shiflet, Thomas N., ed. 1994. Rangeland cover types of the United States. Denver, CO: Society for Range Management. 152 p. 
253. Short, Henry L.; Evans, Wain; Boeker, Erwin L. 1977. The use of natural and modified pinyon pine-juniper woodlands by deer and elk. Journal of Wildlife Management. 41(3): 543-559. 
254. Smith, Arthur D.; Beale, Donald M. 1980. Pronghorn antelope in Utah: some research and observations. Publication No. 80-13. Salt Lake City, UT: Utah Division of Wildlife Resources. 88 p. 
255. Smith, Arthur D.; Hubbard, Richard L. 1954. Preference ratings for winter deer forages from northern Utah ranges based on browsing time and forage consumed. Journal of Range Management. 7: 262-265. 
256. Smith, Dwight R. 1967. Effects of cattle grazing on a ponderosa pine-bunchgrass range in Colorado. Technical Bulletin No. 1371. Washington, DC: U.S. Department of Agriculture, Forest Service. 60 p. 
257. Smith, Dwight Raymond. 1971. Growth responses of true mountain mahogany (Cercocarpus montanus) on four soil types within the Front Range of Colorado. Logan, UT: Utah State University. 206 p. Dissertation. 
258. Springfield, H. W. 1973. Cliffrose and mountainmahogany seeds retain viability 6 years in cold storage. Res. Note RM-236. Fort Collins, CO: U.S. Department of Agriculture, Forest Service, Rocky Mountain Forest and Range Experiment Station. 2 p. 
259. Stahlecker, Dale W.; Kennedy, Patricia L.; Cully, Anne C.; Kuykendall, Charles B. 1989. Breeding bird assemblages in the Rio Grande Wild and Scenic River Recreation Area, New Mexico. The Southwestern Naturalist. 34(4): 487-498. 
260. Stanton, Frank. 1974. Wildlife guidelines for range fire rehabilitation. Tech. Note 6712. Denver, CO: U.S. Department of the Interior, Bureau of Land Management. 90 p. 
261. Steinauer, Gerald A. 1981. A classification of the Cercocarpus montanus, Quercus macrocarpa, Populus deltoides, and Picea glauca habitat types of the Black Hills National Forest. Vermillion, SD: University of South Dakota. 95 p. Thesis. 
262. Stephens, H. A. 1973. Woody plants of the North Central Plains. Lawrence, KS: The University Press of Kansas. 530 p. 
263. Stevens, Richard. 2004. Establishing plants by transplanting and interseeding. In: Monsen, Stephen B.; Stevens, Richard; Shaw, Nancy L., comps. Restoring western ranges and wildlands. Gen. Tech Rep. RMRS-GTR-136-vol-3. Fort Collins, CO: U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station: 739-744. 
264. Stevens, Richard. 2004. Incorporating wildlife habitat needs into restoration and rehabilitation projects. In: Monsen, Stephen B.; Stevens, Richard; Shaw, Nancy L., comps. Restoring western ranges and wildlands. Gen. Tech. Rep. RMRS-GTR-136-vol-1. Fort Collins, CO: U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station: 155-174. 
265. Stevens, Richard. 2004. Management of restored and revegetated sites. In: Monsen, Stephen B.; Stevens, Richard; Shaw, Nancy L., comps. Restoring western ranges and wildlands. Gen. Tech. Rep. RMRS-GTR-136-vol-1. Fort Collins, CO: U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station: 193-198. 
266. Stevens, Richard; Jorgensen, Kent R. 1994. Rangeland species germination through 25 and up to 40 years of warehouse storage. In: Monsen, Stephen B.; Kitchen, Stanley G., comps. Proceedings--ecology and management of annual rangelands; 1992 May 18-22; Boise, ID. Gen. Tech. Rep. INT-GTR-313. Ogden, UT: U.S. Department of Agriculture, Forest Service, Intermountain Research Station: 257-265. 
267. Stevens, Richard; Monsen, Stephen B. 2004. Guidelines for restoration and rehabilitation of principal plant communities. In: Monsen, Stephen B.; Stevens, Richard; Shaw, Nancy L., comps. Restoring western ranges and wildlands. Gen. Tech. Rep. RMRS-GTR-136-vol-1. Fort Collins, CO: U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station: 199-294. 
268. Stickney, Peter F. 1989. FEIS postfire regeneration workshop--April 12: Seral origin of species comprising secondary plant succession in Northern Rocky Mountain forests. 10 p. Unpublished draft on file at: U.S. Department of Agriculture, Forest Service, Intermountain Research Station, Fire Sciences Laboratory, Missoula, MT. 
269. Stormer, Fred A. 1979. Summer habitat of scaled quail in the Texas panhandle. In: Sosebee, Ronald E.; Wright, Henry A.; eds. Research highlights--1979: Noxious brush and weed control; range and wildlife management. Volume 10. Lubbock, TX: Texas Tech University, College of Agricultural Sciences: 58. 
270. Stormer, Fred A. 1984. Night-roosting habitat of scaled quail. Journal of Wildlife Management. 48(1): 191-197. 
271. Stubbendieck, James; Hatch, Stephan L.; Butterfield, Charles H. 1992. North American range plants. 4th ed. Lincoln, NE: University of Nebraska Press. 493 p. 
272. Stuever, Mary C.; Hayden, John S. 1996. Plant associations (habitat types) of the forests and woodlands of Arizona and New Mexico. Final report: Contract R3-95-27. Placitas, NM: Seldom Seen Expeditions, Inc. 520 p. 
273. Stutz, Howard C. 1972. Genetic improvement in crop species as contrasted with possibilities in shrubs. In: McKell, Cyrus M.; Blaisdell, James P.; Goodin, Joe R., eds. Wildland shrubs--their biology and utilization: An international symposium: Proceedings; 1971 July; Logan, UT. Gen. Tech. Rep. INT-1. Ogden, UT: U.S. Department of Agriculture, Forest Service, Intermountain Forest and Range Experiment Station: 139-143. 
274. Stutz, Howard C. 1990. Taxonomy and evolution of Cercocarpus in the western United States. In: Johnson, Kendall L., ed. Proceedings, 5th Utah shrub ecology workshop: The genus Cercocarpus; 1988 July 13-14; Logan, UT. Logan, UT: Utah State University, College of Natural Resources: 15-25. 
275. Swanson, David L.; Palmer, Jeffrey S.; Liknes, Eric T.; Dean, Kurt L. 2000. A breeding population of Virginia's warblers in the southwestern Black Hills of South Dakota. The Southwestern Naturalist. 45(1): 39-44. 
276. Swetnam, Thomas W.; Baisan, Christopher H.; Caprio, Anthony C.; Brown, Peter M. 1992. Fire history in a Mexican oak-pine woodland and adjacent montane conifer gallery forest in southeastern Arizona. In: Ffolliott, Peter F.; Gottfried, Gerald J.; Bennett, Duane A.; Hernandez C., Victor Manuel; Ortega-Rubio, Alfred; Hamre, R. H., tech. coords. Ecology and management of oak and associated woodlands: perspectives in the southwestern United States and northern Mexico: Proceedings; 1992 April 27-30; Sierra Vista, AZ. Gen. Tech. Rep. RM-218. Fort Collins, CO: U.S. Department of Agriculture, Forest Service, Rocky Mountain Forest and Range Experiment Station: 165-173. 
277. Szaro, Robert C. 1981. Bird population responses to converting chaparral to grassland and riparian habitats. The Southwestern Naturalist. 26(3): 251-256. 
278. Taber, Richard D.; Dasmann, Raymond F. 1958. The black-tailed deer of the chaparral: Its life history and management in the north Coast Range of California. Game Bulletin No. 8. Sacramento, CA: State of California, Department of Fish and Game, Game Management Branch. 166 p. 
279. Tande, Gerald F. 1979. Fire history and vegetation pattern of coniferous forests in Jasper National Park, Alberta. Canadian Journal of Botany. 57: 1912-1931. 
280. Tatschl, John L. 1967. Breeding birds of the Sandia Mountains and their ecological distributions. The Condor. 69(5): 479-490. 
281. Thilenius, John F. 1972. Classification of deer habitat in the ponderosa pine forest of the Black Hills, South Dakota. Res. Pap. RM-91. Fort Collins, CO: U.S. Department of Agriculture, Forest Service. 28 p. 
282. Thilenius, John F.; Brown, Gary R.; Medina, Alvin L. 1995. Vegetation on semi-arid rangelands, Cheyenne River Basin, Wyoming. Gen. Tech. Rep. RM-GTR-263. Fort Collins, CO: U.S. Department of Agriculture, Forest Service, Rocky Mountain Forest and Range Experiment Station. 60 p. 
283. Thompson, Robert M. 1999. An example of pinyon-juniper woodland classification in southeastern Utah. In: Monsen, Stephen B.; Stevens, Richard, compilers. Proceedings: ecology and management of pinyon-juniper communities within the Interior West: Sustaining and restoring a diverse ecosystem; 1997 September 15-18; Provo, UT. Proceedings RMRS-P-9. Ogden, UT: U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station: 60-63. 
284. Ticer, Cindy L.; Morrell, Thomas E.; Devos, James C., Jr. 2001. Diurnal bed-site selection of urban-dwelling javelina in Prescott, Arizona. Journal of Wildlife Management. 65(1): 136-140. 
285. Todd, J. W. 1975. Foods of Rocky Mountain bighorn sheep in southern Colorado. Journal of Wildlife Management. 39(1): 108-111. 
286. Tratz, Wallace Michael. 1978. Postfire vegetational recovery, productivity, and herbivore utilization of a chaparral-desert ecotone. Los Angeles, CA: California State University. 133 p. Thesis. 
287. TRW Environmental Safety Systems Inc. 1999. Appendix C: Descriptions of land cover types found within or near Yucca Mountain and the potential transportation corridors and facilities. In: Environmental baseline file for biological resources: B00000000-01717-5700-00009 REV 00. Civilian Radioactive Waste Management System: Management and Operating Contractor--Contract Number DE-AC08-91RW00134, [Online]. North Las Vegas, NV: U.S. Department of Energy, Yucca Mountain Site Characterization Office (Producer). Available: http://www.ymp.gov/documents/biology/appendixc.htm [2000, November 6]. 
288. Turley, Deborah; Roundy, Bruce A.; Walker, Scott C. 2003. Growth and reproductive responses of true mountain mahogany to browsing. Journal of Range Management. 56(6): 591-599. 
289. Turley, Deborah; Roundy, Bruce A.; Walker, Scott C. 2003. True mountain mahogany community and shrub size responses to browsing. Journal of Range Management. 56(6): 600-607. 
290. U.S. Department of Agriculture, Forest Service. 1937. Range plant handbook. Washington, DC. 532 p. 
291. U.S. Department of Agriculture, Natural Resources Conservation Service, Tucson Plant Materials Center. 2001. Commercial sources of conservation plant materials, [Online]. Available: http://plant-materials.nrcs.usda.gov/pubs/azpmsarseedlist0501.pdf [2003, August 25]. 
292. U.S. Department of Agriculture, Natural Resources Conservation Service. 2006. PLANTS database (2006), [Online]. Available: http://plants.usda.gov/. 
293. Urness, P. J.; Neff, D. J.; Watkins, R. K. 1975. Nutritive value of mule deer forages on ponderosa pine summer range in Arizona. Res. Note RM-304. Fort Collins, CO: U.S. Department of Agriculture, Forest Service, Rocky Mountain Forest and Range Experiment Station. 6 p. 
294. Urness, Philip J. 1973. Part II: Chemical analyses and in vitro digestibility of seasonal deer forages. In: Deer nutrition in Arizona chaparral and desert habitats. Special Report No. 3: Federal Aid in Wildlife Restoration Act Project W-78-R. Phoenix, AZ: Arizona Game and Fish Department, Research Division: 39-52. In cooperation with: U.S. Department of Agriculture, Forest Service, Rocky Mountain Forest and Range Experiment Station. 
295. Van Dyne, George M. 1958. Ranges and range plants. [Fort Collins, CO]: [Colorado State University]. Unpublished manuscript on file at: U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory, Missoula, MT. 290 p. 
296. van Wagtendonk, J. W. 1991. Spatial analysis of lightning strikes in Yosemite National Park. In: Andrews, Patricia L.; Potts, Donald F., eds. Proceedings, 11th conference on fire and forest meteorology; 1991 April 16-19; Missoula, MT. SAF Publication 91-04. Bethesda, MD: Society of American Foresters: 605-611. 
297. van Wagtendonk, Jan W. 1987. The role of fire in the Yosemite Wilderness. In: Lucas, Robert C., compiler. Proceedings--national wilderness research conference: issues, state-of-knowledge, future directions; 1985 July 23-26; Fort Collins, CO. Gen. Tech. Rep. INT-220. Ogden, UT: U.S. Department of Agriculture, Forest Service, Intermountain Research Station: 135-177. 
298. Vaughan, Terry A. 1954. Mammals of the San Gabriel Mountains of California. University of Kansas Publications, Museum of Natural History. Lawrence, KS: University of Kansas. 7(9): 513-582. 
299. Vestal, Paul A. 1952. Ethnobotany of the Ramah Navaho. Reports of the Ramah Project: No. 4. Papers of the Peabody Museum of American Archeology and Ethnology. Cambridge, MA: Harvard University. 40(4): 1-94. 
300. Vincent, Dwain W. 1992. The sagebrush/grasslands of the upper Rio Puerco area, New Mexico. Rangelands. 14(5): 268-271. 
301. Vines, Robert A. 1960. Trees, shrubs, and woody vines of the Southwest. Austin, TX: University of Texas Press. 1104 p. 
302. Vogl, Richard J. 1973. Ecology of knobcone pine in the Santa Ana Mountains, California. Ecological Monographs. 43: 125-143. 
303. Vogl, Richard J.; Schorr, Paul K. 1972. Fire and manzanita chaparral in the San Jacinto Mountains, California. Ecology. 53(6): 1179-1188. 
304. Wakimoto, Ronald H. 1977. Chaparral growth and fuel assessment in southern California. In: Mooney, Harold A.; Conrad, C. Eugene, technical coordinators. Proceedings of the symposium on the environmental consequences of fire and fuel management in Mediterranean ecosystems; 1977 August 1-5; Palo Alto, CA. Gen. Tech. Rep. WO-3. Washington, DC: U.S. Department of Agriculture, Forest Service: 412-418. 
305. Wakimoto, Ronald H. 1978. Responses of southern California brushland vegetation to fuel manipulation. Berkely, CA: University of California. 264 p. Dissertation. 
306. Walker, Scott C.; Turley, Deborah. 1999. Characteristics of mountain mahogany (Cercocarpus) species and hybrids in Utah hybrid zone. In: McArthur, E. Durant; Ostler, W. Kent; Wambolt, Carl L., compilers. Proceedings: shrubland ecotones; 1998 August 12-14; Ephraim, UT. Proceedings RMRS-P-11. Ogden, UT: U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station: 32-34. 
307. Wallmo, O. C. 1954. Nesting of Mearns quail in southeastern Arizona. The Condor. 56(3): 125-128. 
308. Wandera, Jackson L. 1990. The basis of browsing tolerance in shrubs of the Intermountain West: growth rates and meristematic potential. Logan, UT: Utah State University. 68 p. Dissertation. 
309. Wangler, Michael J.; Minnich, Richard A. 1996. Fire and succession in pinyon-juniper woodlands of the San Bernardino Mountains, California. Madrono. 43(4): 493-514. 
310. Warner, Thomas E. 1980. Fire history in the yellow pine forest of Kings Canyon National Park. In: Stokes, Marvin A.; Dieterich, John H., technical coordinators. Proceedings of the fire history workshop; 1980 October 20-24; Tucson, AZ. Gen. Tech. Rep. RM-81. Fort Collins, CO: U.S. Department of Agriculture, Forest Service, Rocky Mountain Forest and Range Experiment Station: 89-92. 
311. Warren, Peter L.; Hoy, Marina S.; Hoy, Wilton E. 1992. Vegetation and flora of Fort Bowie National Historic Site, Arizona. Tech. Rep. NPS/WRUA/NRTR-92/43. Tucson, AZ: The University of Arizona, School of Renewable Natural Resources, Cooperative National Park Resources Studies Unit. 78 p. 
312. Waugh, William J. 1990. Stagnation and decadence of Cercocarpus montanus in a southeastern Wyoming big game exclosure. In: Johnson, Kendall L., ed. Proceedings, 5th Utah shrub ecology workshop: The genus Cercocarpus; 1988 July 13-14; Logan, UT. Logan, UT: Utah State University, College of Natural Resources: 89-95. 
313. Weber, William A. 1987. Colorado flora: western slope. Boulder, CO: Colorado Associated University Press. 530 p. 
314. Welch, Bruce L. 1981. Nutritive value of big sagebrush and other shrubs. In: Proceedings--shrub establishment on disturbed arid and semi-arid lands symposium; 1980 December 2-3; Laramie, WY. Laramie, WY: Wyoming Game and Fish Department: 9-22. 
315. Welch, Bruce L. 1989. Nutritive value of shrubs. In: McKell, Cyrus M., ed. The biology and utilization of shrubs. San Diego, CA: Academic Press, Inc: 405-424. 
316. Wells, Michael L.; O'Leary, John F.; Franklin, Janet; Michaelsen, Joel; McKinsey, David E. 2004. Variations in a regional fire regime related to vegetation type in San Diego County, California (USA). Landscape Ecology. 19: 139-152. 
317. Welsh, Stanley L.; Atwood, N. Duane; Goodrich, Sherel; Higgins, Larry C., eds. 1987. A Utah flora. The Great Basin Naturalist Memoir No. 9. Provo, UT: Brigham Young University. 894 p. 
318. West, Neil E.; Tausch, Robin J.; Tueller, Paul T. 1998. A management-oriented classification of pinyon-juniper woodlands of the Great Basin. Gen. Tech. Rep. RMRS-GTR-12. Ogden, UT: U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station. 42 p. 
319. Whittaker, R. H.; Niering, W. A. 1975. Vegetation of the Santa Catalina Mountains, Arizona. V. Biomass, production, and diversity along the elevation gradient. Ecology. 56(4): 771-790. 
320. Wienhold, Brian J.; Klemmedson, James O. 1992. Effect of prescribed fire on nitrogen and phosphorus in Arizona chaparral soil-plant systems. Arid Soil Research and Rehabilitation. 6: 285-296. 
321. Wiggins, Ira L. 1980. Flora of Baja California. Stanford, CA: Stanford University Press. 1025 p. 
322. Williams, Stephen E.; Johnson, Joe D.; Munn, Larry C.; Nieder, Terry. 2004. Edaphic characteristics of nitrogen fixing nodulation (Actinorhizae) by Cercocarpus montanus Raf. and Purshia tridentata (Pursh)DC. In: Hild, Ann L.; Shaw, Nancy L.; Meyer, Susan E.; Booth, D. Terrance; McArthur, E. Durant, compilers. Seed and soil dynamics in shrubland ecosystems: proceedings; 2002 August 12-16; Laramie, WY. Proceedings RMRS-P-31. Fort Collins, CO: U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station: 84-91. 
323. Woodmansee, Robert G. 1969. Natural reproduction of Eurotia lanata, Atriplex canescens, Cercocarpus montanus, and Cowania mexicana in New Mexico. Albuquerque, NM: University of New Mexico. 77 p. Thesis. 
324. Wright, Henry A.; Bailey, Arthur W. 1982. Fire ecology: United States and southern Canada. New York: John Wiley & Sons. 501 p. 
325. Wydeven, Adrian P.; Dahlgren, Robert B. 1983. Food habits of elk in the northern Great Plains. Journal of Wildlife Management. 47(4): 916-923; 1983. 
326. Young, D. Lewis; Bailey, James A. 1975. Effects of fire and mechanical treatment on Cercocarpus montanus and Ribes cereum. Journal of Range Management. 28(6): 495-497. 
327. Young, Daniel Lewis. 1973. First-year effects of fire, clipping, and herbicide on the quantity and quality of current annual growth of two species of shrubs. Fort Collins, CO: Colorado State University. 78 p. Thesis. 
328. Young, James A.; Evans, Raymond A. 1981. Demography and fire history of a western juniper stand. Journal of Range Management. 34(6): 501-505. 
329. Youngblood, Andrew P.; Mauk, Ronald L. 1985. Coniferous forest habitat types of central and southern Utah. Gen. Tech. Rep. INT-187. Ogden, UT: U.S. Department of Agriculture, Forest Service, Intermountain Research Station. 89 p. 
330. Zacek, Joseph C.; Hunter, Harold E.; Bown, T. A.; Ross, Robert L. 1977. Montana grazing guides. [Washington, DC]: U.S. Department of Agriculture, Soil Conservation Service. 12 p. On file with: U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory, Missoula, MT. 
331. Zammit, C.; Zedler, P. H. 1994. Organization of the soil seed bank in mixed chaparral. Vegetatio. 111: 1-16. 
332. Zedler, Paul H. 1977. Life history attributes of plants and the fire cycle: a case study in chaparral dominated by Cupressus forbesii. In: Mooney, Harold A.; Conrad, C. Eugene, technical coordinators. Symposium on the environmental consequences of fire and fuel management on Mediterranean ecosystems: Proceedings; 1977 August 1-5; Palo Alto, CA. Gen. Tech. Rep. WO-3. Washington, DC: U.S. Department of Agriculture, Forest Service: 451-458. 
333. Zedler, Paul H. 1981. Vegetation change in chaparral and desert communities in San Diego County, California. In: West, D. C.; Shugart, H. H.; Botkin, D. B., eds. Forest succession: concepts and application. New York: Springer-Verlag: 406-430. 
334. Zedler, Paul H. 1995. Fire frequency in southern California shrublands: biological effects and management options. In: Keeley, Jon F.; Scott, Tom, eds. Brushfires in California: ecology and resource management: Proceedings; 1994 May 6-7; Irvine, CA. Fairfield, WA: International Association of Wildland Fire: 101-112. 
335. Zedler, Paul H.; Gautier, Clayton R.; McMaster, Gregory S. 1983. Vegetation change in response to extreme events: the effect of a short interval between fires in California chaparral and coastal scrub. Ecology. 64(4): 809-818. 
336. Zigmond, Maurice L. 1981. Kawaisu ethnobotany. Salt Lake City, UT: University of Utah Press. 102 p. 
FEIS Home Page