Index of Species Information

SPECIES:  Dichanthelium acuminatum
Photo by James H. Miller & Ted Bodner, Southern Weed Science Society,


SPECIES: Dichanthelium acuminatum
AUTHORSHIP AND CITATION : Walsh, Roberta A. 1995. Dichanthelium acuminatum. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: []. Revisions : On 9 February 2016, the common name of this species was changed from: hairy panicgrass to: tapered rosette grass. Additional synonyms and the following citations were added: [44,53] ABBREVIATION : DICACU SYNONYMS : Dichanthelium acuminatum (Sw.) Gould & C.A. Clark subsp. acuminatum Dichanthelium acuminatum subsp. columbianum (Scribn.) Freckman & LeLong, panic du District of Columbia Dichanthelium acuminatum subsp. fasiculatum (Torr.) Freckman & LeLong Dichanthelium acuminatum subsp. implicatum (Scribn.) Freckman & LeLong Dichanthelium acuminatum subsp. lindheimeri (Nash) Freckman & LeLong Dichanthelium acuminatum subsp. sericeum (Schmoll) Freckman & LeLong Dichanthelium acuminatum subsp. thermale (Bol.) Freckman & LeLong [53] Dichanthelium acuminatum var. implicatum (Scribn.) Gould & Clark [20,26] = Dichanthelium acuminatum var. fasciculatum (Torr.) Freckmann [44] Panicum lanuginosum Ell. [15,17,20,21,23,24,36,40,52] NRCS PLANT CODE [53] : DIAC2 DIACF DIACL DIACS DIACT2 DIACT COMMON NAMES : tapered rosette grass green panicgrass hairy panicgrass panic laineux woolly panicum TAXONOMY : The currently accepted scientific name of tapered rosette grass is Dichanthelium acuminatum (Sw.) Gould & C.A. Clark (Poaceae) [4,20,26]. The following varieties are generally accepted [20,26,44]: Dichanthelium acuminatum var. acuminatum Dichanthelium acuminatum var. fasciculatum (Torr.) Freckmann, western panicgrass Dichanthelium acuminatum var. lindheimeri (Nash) Gould & Clark, Lindheimer panicgrass Dichanthelium acuminatum var. sericeum (Schmoll) Freckmann, Pacific panicgrass Dichanthelium acuminatum var. thermale (Bol.) Freckmann, Geyser's panicgrass Dichanthelium acuminatum var. thurowii (Scribn. & J. G. Smith) Gould & Clark, Thurow's panicgrass The above taxa have received many treatments. In 1910, Hitchcock and Chase placed some Panicum species into the subgenus Dichanthelium, which was elevated to generic status by Gould in 1974. Gould and Clark extensively revised Dichanthelium in 1978 [20]. Freckmann produced a revision in 1981, reassigning some former D. acuminatum varieties to species level [15]. LIFE FORM : Graminoid FEDERAL LEGAL STATUS : No special status OTHER STATUS : NO-ENTRY


SPECIES: Dichanthelium acuminatum
GENERAL DISTRIBUTION : Tapered rosette grass is widespread in the Americas. It occurs throughout the United States and in Canada from Newfoundland and Nova Scotia west to British Columbia. Its distribution extends from eastern Mexico south to northern South America and it is found in the Antilles [8,14,17,21,22,23,24,27,29,40]. ECOSYSTEMS : Tapered rosette grass occurs in most ecosystems. STATES : AL AK AZ AR CA CO CT DE FL GA HI ID IL IN IA KS KY LA ME MD MA MI MN MS MO MT NE NV NH NJ NM NY NC ND OH OK OR PA RI SC SD TN TX UT VT VA WA WV WI WY DC PR VI AB BC MB NB NF NS ON PE PQ SK MEXICO BLM PHYSIOGRAPHIC REGIONS : 1 Northern Pacific Border 2 Cascade Mountains 3 Southern Pacific Border 4 Sierra Mountains 5 Columbia Plateau 6 Upper Basin and Range 7 Lower Basin and Range 8 Northern Rocky Mountains 9 Middle Rocky Mountains 10 Wyoming Basin 11 Southern Rocky Mountains 12 Colorado Plateau 13 Rocky Mountain Piedmont 14 Great Plains 15 Black Hills Uplift 16 Upper Missouri Basin and Broken Lands KUCHLER PLANT ASSOCIATIONS : Tapered rosette grass occurs in most Kuchler Plant Associations. SAF COVER TYPES : Tapered rosette grass occurs in most SAF Cover Types. SRM (RANGELAND) COVER TYPES : Tapered rosette grass occurs in most SRM Cover Types. HABITAT TYPES AND PLANT COMMUNITIES : Tapered rosette grass occurs in many habitat types and within diverse plant communities. On dry prairie ridges of central Wisconsin sand plains, associates of tapered rosette grass include little bluestem (Schizachyrium scoparium), skyblue aster (Aster oolentangiensis), heath aster (A. ericoides), forked threeawn (Aristida basiramea), parasol sedge (Carex umbellata), poverty oatgrass, prairie junegrass (Koeleria macrantha), tall gayfeather (Liatris aspera), and Indiangrass (Sorghastrum nutans) [50]. In central and northwestern Wisconsin, tapered rosette grass is found on sandy outwashes and former lake bottoms. Associates include ragweed (Ambrosia psilostachya), clasping milkweed (Asclepias amplexicaulis), longbranch frostweed (Helianthemum canadense), Kalm's hawkweed (Hieracium kalmii), hairy hawkweed (H. longipilum), racemed milkwort (Polygala polygama), and Virginia tephrosia (Tephrosia virginiana) [46]. On the west slope of the Sierra Nevada in Placer County, California, tapered rosette grass occurs in grazed meadows which follow natural drainages. Associates include Thurber's bentgrass (Agrostis thurberiana), other bentgrasses (Agrostis spp.), California oatgrass (Danthonia californica), pullup muhly (Muhlenbergia filiformis), sedges (Carex spp.), rushes (Juncus spp.), woodrushes (Luzula spp.), white clover (Trifolium repens), seep monkeyflower (Mimulus guttatus), primrose monkeyflower (M. primuloides), trailing St. Johnswort (Hypericum anagalloides), and violets (Viola spp.) [1]. In the Adirondacks of New York, tapered rosette grass associates include Pickering's reedgrass (Calamagrostis pickeringii), bluejoint reedgrass (C. canadensis), northern reedgrass (C. stricta ssp. inexpansa), sedges, poverty oatgrass (Danthonia spicata), threeway sedge (Dulichium arundinaceum), rattlesnake mannagrass (Glyceria canadensis), narrowpanicle rush (Juncus brevicaudatus), brownfruit rush (J. pelocarpus), bog muhly (Muhlenbergia uniflora), whitebeaked rush (Rhynchospora alba), and low bulrush (Scirpus cespitosus) [51].


SPECIES: Dichanthelium acuminatum
IMPORTANCE TO LIVESTOCK AND WILDLIFE : NO-ENTRY PALATABILITY : NO-ENTRY NUTRITIONAL VALUE : NO-ENTRY COVER VALUE : NO-ENTRY VALUE FOR REHABILITATION OF DISTURBED SITES : Tapered rosette grass is useful as a soil stabilizer and for restoring degraded areas [23]. However, in some locations it declines in importance after a few years as other species increase in cover. By 1976, tapered rosette grass was thriving along a section of highway in Peoria, Illinois, that had been planted to prairie vegetation in November 1966. Prairie seeds had been broadcast over hard-packed clay soil that had been loosened [5]. Prairie was reestablished at Pea Ridge National Military Park, Arkansas. The original prairie had been turned to agricultural use. The site was plowed in early June 1975, and a seed mixture (not including woolly panicum) was hand-broadcast on each plot. Vegetation was sampled in early fall of 1975 and in late spring and early fall for 4 years thereafter. Tapered rosette grass established in the fall of the first year and persisted for the 5 years of the study. However, it was prominent only in the 2-year-old plots. Seeded prairie grasses in the restoration plots have increased, and other grasses including tapered rosette grass have decreased. Mean percent cover of tapered rosette grass on 1- to 5-year-old plots based on spring (S) and fall (F) sampling was as follows [6]: Plot Age in Years 1 2 3 4 5 F S F S F S F S F 0.5 4.8 5.0 2.1 1.6 0.5 0.0 1.0 0.0 Tapered rosette grass was a component of 4-year-old and 8-year-old reestablished prairies in southwestern Ohio [31]. OTHER USES AND VALUES : NO-ENTRY OTHER MANAGEMENT CONSIDERATIONS : Tapered rosette grass does not respond uniformly to grazing. In the Sierra Nevada in Placer County, California, tapered rosette grass was present in meadows which were grazed May 30 to September 30 each year from 1979 to 1986. Utilization of meadow species averaged 61 percent over the 10 years, but increased to more than 80 percent after 1985. Tapered rosette grass increased from 0.8 to 2.1 percent relative cover over that time [1]. In a 12-year study of grazing intensity in central Louisiana, the frequency of several species, including tapered rosette grass, was greater on ungrazed than on moderately or heavily grazed range. However, differences owing to grazing intensity were considered "small and relatively unimportant" [11].


SPECIES: Dichanthelium acuminatum
GENERAL BOTANICAL CHARACTERISTICS : Tapered rosette grass is a native, perennial bunchgrass [8]. It is polymorphic, having early and late season forms of the same plant. Vernal culms are tufted, usually unbranched and radiating from the base, and 6 to 32 inches (15-80 cm) tall [14,17,36]. Early leaf blades are 1.6 to 3.9 inches (4-10 cm) long [21,22]; panicles are pyramidal and 1.2 to 4.7 inches (3-12 cm) long [14,21,24]. Autumnal culms are decumbent and much-branched, mostly from the middle nodes; they produce dense fascicles of leaves and inflorescences [19,24]. Fall culm leaf blades are shorter, narrower, and more crowded than early blades. Winter rosette blades are also short, but are relatively broad. Late panicles are reduced and often partly enclosed in leaf sheaths [21]. Spikelets are awnless [4]; the fruit is a caryopsis [36]. RAUNKIAER LIFE FORM : Hemicryptophyte REGENERATION PROCESSES : Tapered rosette grass reproduces by seeds; it does not have rhizomes or stolons [17,22]. Vernal panicles are open and produce chasmogamous flowers. Autumnal panicles are reduced [20] and often are partly enclosed in the leaf sheaths [17]; these enclosed flowers are self-pollinated [21]. Tapered rosette grass seeds can be found in the seedbank, but presence of tapered rosette grass vegetation does not necessarily correspond to presence in the seedbank. In southwestern Ohio, five unburned old-field-deciduous forest sites were surveyed for vegetational and seedbank presence. Tapered rosette grass was present as vegetation and in the seedbank at two sites, present only as vegetation at two sites, and not present in either form at one site [37,47,48]. In Harvard Forest at Petersham, Massachusetts, tapered rosette grass was a widespread seedbank component in old fields and white pine (Pinus strobus) stands. It was not a component of the herbaceous ground cover at any site [30]. SITE CHARACTERISTICS : Tapered rosette grass occurs in a wide variety of habitats. It is found in woodlands [19,24] woodland borders, and recently cleared woodland areas. It occurs on prairies [22], dunes, seashores [17], and along open roadsides [20]. It grows along the banks of rivers and lakes [29] and in marshes, swamps and swales [20]. It is found along the shorelines of bogs [39], pondcypress (Taxodium distichum) ponds [4], and around hot springs [9]. Tapered rosette grass grows on many soil types. It occurs on heavy, wet soils [39], silty loam [2,11,48], sandy loam with high organic content [12], moist to dry sand [14,20,24,34,46,49], and gravelly, often calcareous soils [49]. Soils are often acidic [12]. In southern Quebec near Montreal, tapered rosette grass grows on fine loamy sand with pH 6.1; the soil is low in magnesium and very low in calcium and nitrogen [43]. Tapered rosette grass has been reported from the following elevations: Feet Meters Arizona 4,000-6,500 1,219-1,981 [27] California < 8,531 < 2,600 [23] Colorado 3,500-5,500 1,067-1,676 [8,22] New York < 5,348 < 1,630 [51] Utah 3,510-7,317 1,070-2,230 [52] Mexico 98-9,843 30-3,000 [19] SUCCESSIONAL STATUS : Tapered rosette grass appears in early secondary succession. It does not appear in old-growth forests, but is found in stands of intermediate age. In southwestern Ohio, tapered rosette grass occurred in all but the oldest of a chronosequence of five unburned old-field-deciduous forest sites; the sites were surveyed nine times during the 1980 growing season. Site 1 was an old field abandoned for 2 years and dominated by red clover (Trifolium pratense). Site 2 was an old field abandoned for 10 years and dominated by Canada goldenrod (Solidago canadensis) and meadow fescue (Festuca pratensis). Site 3 was an old field abandoned for 50 years and dominated by Canada goldenrod, with white ash (Fraxinus americana) and black cherry (Prunus serotina) providing about 30 percent canopy cover. Site 4 was an old field abandoned for 90 years and dominated by a forest of sugar maple (Acer saccharum) and slippery elm (Ulmus rubra). Site 5 was an old-growth American beech (Fagus grandifolia) and sugar maple dominated forest which was over 200 years old. Tapered rosette grass was present but not common at site 1; its seeds were present in the seedbank. It was present in "disturbed" vegetation at site 2; its seeds were not found in the seedbank. Tapered rosette grass was a common species in "disturbed" vegetation at site 3; its seeds were found in the seedbank. It was present but not common at site 4; its seeds were not found in the seedbank. Tapered rosette grass was not present nor found in the seedbank at site 5 [37,47,78]. In upland oak-pine forest sites in the New Jersey Pine Barrens, woolly panicum did not occur in the flora but appeared to persist in the seedbank. Its persistence was attributed to infrequent local disturbance [32]. Tapered rosette grass was found in marshes of a Connecticut River oxbow in western Massachusetts in 1984, though it had not been present in 1974. Tapered rosette grass was found only in the highest and driest areas of the marshes, at low cover (<0.5%) and frequency (1%). During high water, much of the marsh area is flooded. The appearance and disappearance of tapered rosette grass was probably due to variations in water level during the study period [25]. SEASONAL DEVELOPMENT : Tapered rosette grass produces a basal rosette during autumn and winter. Primary inflorescences develop and bloom on elongated unbranched culms that grow from the rosettes early in spring. Late spring, summer, and autumn growth of the same culms produces fascicles of short, branched and rebranched lateral shoots with greatly reduced leaves and lateral inflorescences which also bloom. Tapered rosette grass thus has two main flowering periods [20]. Tapered rosette grass flowering times are: California June-August [35] Florida May-September [4] Illinois May-October [34] New York June-July [10] West Virginia June-September [42] Eastern United States May-November [14] Great Plains May-June [21] through fall New England June-September [40] Southeastern United States May-August [36] Southern United States begins [20] February-March


SPECIES: Dichanthelium acuminatum
FIRE ECOLOGY OR ADAPTATIONS : Tapered rosette grass has basal buds [21] which may sprout after aerial portions are burned. If thick tufts form, they may protect the basal buds from fire damage. FIRE REGIMES : Find fire regime information for the plant communities in which this species may occur by entering the species name in the FEIS home page under "Find Fire Regimes". POSTFIRE REGENERATION STRATEGY : Tussock graminoid Ground residual colonizer (on-site, initial community)


SPECIES: Dichanthelium acuminatum
IMMEDIATE FIRE EFFECT ON PLANT : Tapered rosette grass culms and leaves are probably killed by fire. DISCUSSION AND QUALIFICATION OF FIRE EFFECT : NO-ENTRY PLANT RESPONSE TO FIRE : Tapered rosette grass presence in response to fire varies with frequency and severity of fire. Tapered rosette grass occurs with greater frequency following annual fires than following less frequent fires. It has lowest frequency where there are no fires [7,12,31,33]. However, if the fire is severe enough to damage buds and roots, tapered rosette grass presence declines [2]. Fire reduces tapered rosette grass flowering [2,18]. DISCUSSION AND QUALIFICATION OF PLANT RESPONSE : Tapered rosette grass was subjected to annual or periodic surface fires in upland oak forests in Franklin County, Tennessee. Data were collected each September from 1965 to 1970. Burning on annual burn plots began in 1963; periodic burn plots were treated in 1964 and 1969. All fires were in late winter. Tapered rosette grass occurred with greatest frequency on annually burned plots, with intermediate frequency on periodically burned plots, and with lowest frequency on unburned plots [7]. A 90-year-old red pine (Pinus resinosa) and white pine stand in Chalk River, Ontario, was surveyed for vegetation in late August of 1969, 1970, 1971, and 1972. Prior to burning, the site was dominated by shrubs and balsam fir (Abies balsamea) saplings. Fires were conducted on June 15 of 1970 and 1971. Total reduction in depth of the organic layer was approximately 30 percent. Fuel consumption of the forest floor was confined to the litter layer; the duff was almost undisturbed. Postfire cover was dominated by herbaceous species. Tapered rosette grass was absent from prefire permanent plots but occurred in the plots in postfire years 1 to 4. However, it did not become important in terms of density or biomass during the time of the study [33]. Tapered rosette grass was a component of a pitcher plant (Sarracenia alata) bog in Harrison County, Mississippi, that had been burned annually in winter over the previous 7 years [12]. In southwestern Ohio, tapered rosette grass was a component of an 8-year-old prairie which was established in 1979 and burned in 1983 [31]. In southern Illinois grassland, tapered rosette grass declined sharply, from 18,415 to 1,080 flowering culms per acre (45,486-2,668 culms/ha), following fire. Plots had been undisturbed from 1950 to 1969. Plots were sampled in August 1969. Burning was conducted December 1, 1969. The fire was moderate, resulting in nearly complete consumption of the heavy litter accumulation from years of fire protection. Postfire sampling of burned and control plots was done from June through September 1970. Tapered rosette grass had the greatest decrease in frequency following burning of any species sampled, probably because of damage to its buds and roots. Response of tapered rosette grass to the fire was [2]: Frequencies (%) Control Burn 1969 1970 1969 1970 24.6 26.3 42.5 0.8 In Illinois, tapered rosette grass showed a 94 percent decrease in flowering following a spring fire [18]. FIRE MANAGEMENT CONSIDERATIONS : NO-ENTRY


SPECIES: Dichanthelium acuminatum
REFERENCES : 1. Allen, Barbara H. 1989. Ten years of change in Sierran stringer meadows: an evaluation of range condition models. In: Abell, Dana L., technical coordinator. Proceedings of the California riparian systems conference: Protection, management, and restoration for the 1990's; 1988 September 22-24; Davis, CA. Gen. Tech. Rep. PSW-110. Berkeley, CA: U.S. Department of Agriculture, Forest Service, Pacific Southwest Forest and Range Experiment Station: 102-108. [13513] 2. Anderson, Roger C.; Van Valkenburg, Charles. 1977. Response of a southern Illinois grassland community to burning. Transactions, Illinois State Academy of Science. 69(4): 399-414. [19481] 3. Bernard, Stephen R.; Brown, Kenneth F. 1977. Distribution of mammals, reptiles, and amphibians by BLM physiographic regions and A.W. Kuchler's associations for the eleven western states. Tech. Note 301. Denver, CO: U.S. Department of the Interior, Bureau of Land Management. 169 p. [434] 4. Clewell, Andre F. 1985. Guide to the vascular plants of the Florida Panhandle. Tallahassee, FL: Florida State University Press. 605 p. [13124] 5. Cull, Margaret Irene. 1978. Establishing prairie vegetation along highways in the Peoria area. In: Glenn-Lewin, David C.; Landers, Roger Q., Jr., eds. Proceedings, 5th Midwest prairie conference; 1976 August 22-24; Ames, IA. Ames, IA: Iowa State University: 172-177. [3378] 6. Dale, Edward D., Jr.; Smith, Thomas C. 1983. Changes in vegetation on a restored prairie at Pea Ridge National Military Park, Arkansas. In: Kucera, Clair L., ed. Proceedings, 7th North American prairie conference; 1980 August 4-6; Springfield, MO. Columbia, MO: University of Missouri: 237-241. [3224] 7. DeSelm, H. R.; Clebsch, E. E. C.; Nichols, G. M.; Thor, E. 1974. Response of herbs, shrubs and tree sprouts in prescribed-burn hardwoods in Tennessee. In: Proceedings, annual Tall Timbers fire ecology conference; 1973 March 22-23; Tallahassee, FL. No. 13. Tallahassee, FL: Tall Timbers Research Station: 331-344. [19011] 8. Dittberner, Phillip L.; Olson, Michael R. 1983. The plant information network (PIN) data base: Colorado, Montana, North Dakota, Utah, and Wyoming. FWS/OBS-83/86. Washington, DC: U.S. Department of the Interior, Fish and Wildlife Service. 786 p. [806] 9. Dorn, Robert D. 1988. Vascular plants of Wyoming. Cheyenne, WY: Mountain West Publishing. 340 p. [6129] 10. Dowhan, Joseph J.; Rozsa, Ron. 1989. Flora of Fire Island, Suffolk County, New York. Bulletin of the Torrey Botanical Club. 116(3): 265-282. [22041] 11. Duvall, V. L.; Linnartz, N. E. 1967. Influences of grazing and fire on vegetation and soil of longleaf pine - bluestem range. Journal of Range Management. 20: 241-247. [7623] 12. Eleuterius, L. N.; Jones, S. B., Jr. 1969. A floristic and ecological study of pitcher plant bogs in south Mississippi. Rhodora. 71: 29-34. [12333] 13. Eyre, F. H., ed. 1980. Forest cover types of the United States and Canada. Washington, DC: Society of American Foresters. 148 p. [905] 14. Fernald, Merritt Lyndon. 1950. Gray's manual of botany. [Corrections supplied by R. C. Rollins]. Portland, OR: Dioscorides Press. 1632 p. (Dudley, Theodore R., gen. ed.; Biosystematics, Floristic & Phylogeny Series; vol. 2). [14935] 15. Freckmann, Robert W. 1981. Realignments in the Dichanthelium acuminatum complex (Poaceae). Phytologia. 48(1): 99-110. [23994] 16. Garrison, George A.; Bjugstad, Ardell J.; Duncan, Don A.; [and others]. 1977. Vegetation and environmental features of forest and range ecosystems. Agric. Handb. 475. Washington, DC: U.S. Department of Agriculture, Forest Service. 68 p. [998] 17. Gleason, Henry A.; Cronquist, Arthur. 1991. Manual of vascular plants of northeastern United States and adjacent Canada. 2nd ed. New York: New York Botanical Garden. 910 p. [20329] 18. Glenn-Lewin, David C.; Johnson, Louise A.; Jurik, Thomas W.; [and others]. 1990. Fire in central North American grasslands: vegetative reproduction, seed germination, and seedling establishment. In: Collins, Scott L.; Wallace, Linda L., eds. Fire in North American tallgrass prairies. Norman, OK: University of Oklahoma Press: 28-45. [14194] 19. Gould, Frank W. 1980. The Mexican species of Dichanthelium (Poaceae). Brittonia. 32(3): 353-364. [23992] 20. Gould, Frank W.; Clark, Carolyn A. 1978. Dichanthelium (Poaceae) in the United States and Canada. Annals of the Missouri Botanical Garden. 65: 1088-1132. [23993] 21. Great Plains Flora Association. 1986. Flora of the Great Plains. Lawrence, KS: University Press of Kansas. 1392 p. [1603] 22. Harrington, H. D. 1964. Manual of the plants of Colorado. 2d ed. Chicago: The Swallow Press Inc. 666 p. [6851] 23. Hickman, James C., ed. 1993. The Jepson manual: Higher plants of California. Berkeley, CA: University of California Press. 1400 p. [21992] 24. Hitchcock, A. S. 1951. Manual of the grasses of the United States. Misc. Publ. No. 200. Washington, DC: U.S. Department of Agriculture, Agricultural Research Administration. 1051 p. [2nd edition revised by Agnes Chase in two volumes. New York: Dover Publications, Inc.]. [1165] 25. Holland, Marjorie M.; Burk, C. John. 1990. The marsh vegetation of three Connecticut River oxbows: a ten-year comparison. Rhodora. 92(871): 166-204. [14521] 26. Kartesz, John T. 1994. A synonymized checklist of the vascular flora of the United States, Canada, and Greenland. Volume II--thesaurus. 2nd ed. Portland, OR: Timber Press. 816 p. [23878] 27. Kearney, Thomas H.; Peebles, Robert H.; Howell, John Thomas; McClintock, Elizabeth. 1960. Arizona flora. 2d ed. Berkeley, CA: University of California Press. 1085 p. [6563] 28. Kuchler, A. W. 1964. Manual to accompany the map of potential vegetation of the conterminous United States. Special Publication No. 36. New York: American Geographical Society. 77 p. [1384] 29. Lackschewitz, Klaus. 1991. Vascular plants of west-central Montana--identification guidebook. Gen. Tech. Rep. INT-227. Ogden, UT: U.S. Department of Agriculture, Forest Service, Intermountain Research Station. 648 p. [13798] 30. Livingston, R. B.; Allessio, Mary L. 1968. Buried viable seed in successional field and forest stands, Harvard Forest, Massachusetts. Bulletin of the Torrey Botanical Club. 95(1): 58-69. [3377] 31. Lutz, Kimberly A. 1989. Prairie establishment in southwestern Ohio. In: Bragg, Thomas B.; Stubbendieck, James, eds. Prairie pioneers: ecology, history and culture: Proceedings, 11th North American prairie conference; 1988 August 7-11; Lincoln, NE. Lincoln, NE: University of Nebraska: 95-99. [14025] 32. Matlack, Glenn R.; Good, Ralph E. 1990. Spatial heterogeneity in the soil seed bank of a mature coastal plain forest. Bulletin of the Torrey Botanical Club. 117(2): 143-152. [22905] 33. Methven, Ian R. 1973. Fire, succession and community structure in a red and white pine stand. Information Report PS-X-43. Chalk River, ON: Environment Canada, Forestry Service, Petawawa Forest Experiment Station. 18 p. [18601] 34. Mohlenbrock, Robert H. 1986. (Revised edition). Guide to the vascular flora of Illinois. Carbondale, IL: Southern Illinois University Press. 507 p. [17383] 35. Munz, Philip A. 1973. A California flora and supplement. Berkeley, CA: University of California Press. 1905 p. [6155] 36. Radford, Albert E.; Ahles, Harry E.; Bell, C. Ritchie. 1968. Manual of the vascular flora of the Carolinas. Chapel Hill, NC: The University of North Carolina Press. 1183 p. [7606] 37. Roberts, Teresa L.; Vankat, John L. 1991. Floristics of a chronosequence corresponding to old field-deciduous forest succession in southwestern Ohio. II. Seed banks. Bulletin of the Torrey Botanical Club. 118(4): 377-384. [17753] 38. Raunkiaer, C. 1934. The life forms of plants and statistical plant geography. Oxford: Clarendon Press. 632 p. [2843] 39. Roland, A. E.; Smith, E. C. 1969. The flora of Nova Scotia. Halifax, NS: Nova Scotia Museum. 746 p. [13158] 40. Seymour, Frank Conkling. 1982. The flora of New England. 2d ed. Phytologia Memoirs 5. Plainfield, NJ: Harold N. Moldenke and Alma L. Moldenke. 611 p. [7604] 41. Stickney, Peter F. 1989. Seral origin of species originating in northern Rocky Mountain forests. Unpublished draft on file at: U.S. Department of Agriculture, Forest Service, Intermountain Research Station, Fire Sciences Laboratory, Missoula, MT; RWU 4403 files. 7 p. [20090] 42. Strausbaugh, P. D.; Core, Earl L. 1977. Flora of West Virginia. 2nd ed. Morgantown, WV: Seneca Books, Inc. 1079 p. [23213] 43. Swales, Dorothy E. 1975. An unusual habitat for Drosera rotundifolia L., its over-wintering state, and vegetative reproduction. Canadian Field-Naturalist. 89(2): 143-147. [22335] 44. U.S. Department of Agriculture, Natural Resources Conservation Service. 2016. PLANTS Database, [Online]. Available: [34262] 45. U.S. Department of the Interior, National Biological Survey. [n.d.]. NP Flora [Data base]. Davis, CA: U.S. Department of the Interior, National Biological Survey. [23119] 46. Umbanhowar, Charles E., Jr. 1992. Reanalysis of the Wisconsin prairie continuum. American Midland Naturalist. 127(2): 268-275. [18194] 47. Vankat, John L.; Carson, Walter P. 1991. Floristics of a chronosequence corresponding to old field-deciduous forest success. in southwestern Ohio. III. Post-disturbance vegetation. Bulletin of the Torrey Botanical Club. 118(4): 385-391. [17755] 48. Vankat, John L.; Snyder, Gary W. 1991. Floristics of a chronosequence corresponding to old field-diciduous forest succession in southwestern Ohio. I. Undisturbed vegetation. Bulletin of the Torrey Botanical Club. 118(4): 365-376. [18758] 49. Voss, Edward G. 1972. Michigan flora. Part I. Gymnosperms and monocots. Bloomfield Hills, MI: Cranbrook Institute of Science; Ann Arbor, MI: University of Michigan Herbarium. 488 p. [11471] 50. Zedler, Joy; Loucks, Orie L. 1969. Differential burning response of Poa pratensis fields and Andropogon scoparius prairies in central Wisconsin. American Midland Naturalist. 81(2): 341-352. [14795] 51. Zika, Peter F.; Jenkins, Jerry C. 1992. Contributions to the flora of the Adirondacks, New York. Bulletin of the Torrey Botanical Club. 119(4): 442-445. [20903] 52. Welsh, Stanley L.; Atwood, N. Duane; Goodrich, Sherel; Higgins, Larry C., eds. 1987. A Utah flora. Great Basin Naturalist Memoir No. 9. Provo, UT: Brigham Young University. 894 p. [2944] 53. Barkworth, Mary E.; Capels, Kathleen M.; Long, Sandy; Piep, Michael B., eds. 2003. Flora of North America north of Mexico. Volume 25: Magnoliophyta: Commelinidae (in part): Poaceae, part 2. New York: Oxford University Press. 814 p. [68091]