Greg Hayward Return to National Staff Page


Wildlife Ecologist Rocky Mountain Region


Rocky Mountain Regional OfficeVoice: (303) 275-5022

740 Simms Street FAX: (303 275-5075

Golden, CO 80401E-mail:




Greg joined the Rocky Mountain Region as the Wildlife Ecologist in 1999 after 19 years as a research ecologist. Experiences working with Oz Garton developing approaches to wildlife monitoring, exploring community ecology, and studying the relationship of vertebrates to vegetation dynamics provided a foundation for his interests in the interaction between management and research. A post-doc with John Wiens examining the effects of the Exxon Valdez oil spill on seabirds introduced Greg to formal approaches to landscape ecology. He took his current position after serving as an Assistant Professor at the University of Wyoming where he won the department's Outstanding Teaching Award while conducting research on topics ranging from the genetic structure of boreal owls in North America to modeling the interaction of cutthroat and lake trout in Yellowstone Lake, to developing monitoring systems for the Amur Tiger in the Russian Far East. Greg maintains a position on the faculty at the University of Wyoming where he teaches landscape ecology and continues to work with his university colleagues.




BS Colorado State University, 1979
MS University of Idaho, 1983
PHD University of Idaho, 1989


Selected Publications


Stephens, P.A., Buskirk, S.W., Hayward, G.D. & Martínez del Rio, C. 2005 Information theory and hypothesis testing: a call for pluralism. Journal of Applied Ecology, 42, 4-12


Marni E. Koopman, David B. McDonald, Gregory D. Hayward, Katrine Eldegard, Geir A. Sonerud and Sergey G. Sermach. 2005. Genetic similarity among Eurasian subspecies of boreal owls Aegolius funereus. Journal of Avian Biology 36: 1-5.


Keinath, D. A., and G. D. Hayward. 2003. Red-backed vole (Clethrionomys gapperi) response to disturbance in subalpine forests: use of regenerating patches. J. Mammalogy 84:956-966.


Hayward, G. D. D. Miquelle, E. N. Smirnov, and C. Nations. 2002 Monitoring Amur tiger populations: characteristics of track surveys in snow. Wildlife Society Bulletin. 30:1150-1159.


Stapp, P. and G. D. Hayward. 2002. Effects of an introduced piscivore on native trout: Insights from a demographic model. Biological Invasions 4:299-316


Wiens, J. A., T. O. Crist, R. H. Day, S. M. Murphy, and G. D. Hayward. 2001. A canonical correspondence analysis of the effects of the Exxon Valdez oil spill on marine birds. Ecological Applications. 11:828-839.

Hayward, G. D., J. F. Shogren, and J. Tschirhart. 2001 The nature of endangered species protection. in: J. Shogren and J. Tschirhart, eds. Protecting endangered species in the United States: Biological needs, political realities, economic choices. Cambridge University Press, New York.


Hayward, G. D., S. H. Henry, and L. Ruggiero. 1999. Response of red-backed voles to recent patch cutting in subalpine forest. Conservation Biology 13:168-176.


Hayward, G. D. and G. C. Iverson. 1998. Long-term trend in marbled murrelets in southeast Alaska based on Christmas Bird Counts. Northwest Science 72:170-179.


Squires, J. and G. D. Hayward. 1998. The role of sensitive species in avian conservation. Pages 155-179. in J. M. Marzluff and R. Sallabanks. eds. Avian Conservation: Research and management. Island Press, New York, N.Y.


Hayward, G. D. 1997. Forest management and conservation of boreal owls in North America. J. Raptor Research 31:114-124.


Rosentretter, R., G. D. Hayward, and M. Wicklow-Howard. 1997. Northern flying squirrel seasonal food habits in the interior conifer forests of central Idaho. Northwest Science 71:97-102.


Day, R. H., S. M. Murphy, J. A. Wiens, G. D. Hayward, E. J. Harner, and L. N. Smith. 1997. Effects of the Exxon Valdez oil spill on habitat use by birds in Prince William Sound, Alaska. Ecological Applications 7:593-613.


Iverson, C, G. D. Hayward, K. Titus, G. Degayner, R. Lowell, C. Crocker-Bedford, P. Schempf, and J. Lindell. 1996. Conservation assessment for northern goshawk in southeast Alaska. U. S. Forest Service, Gen Tech Rep. PNW-387.


Hayward, G. D. and P. H. Hayward. 1995. Relative abundance and habitat associations of small mammals in Chamberlain Basin, central Idaho. Northwest Science. 69:114-125.


Ruggiero, L. F., G. D. Hayward, and J. R. Squires. 1994. Viability analysis in biological evaluations: Concepts of population viability analysis, biological population, and ecological scale. Conservation Biology 8:364-372.


Hayward, G. D. and J. Verner (eds). 1994. Flammulated, boreal, and great gray owls in the United States: A technical conservation assessment. U. S. Forest Service, Gen Tech Rep. RM-253. 214 pp.


Hayward, G. D., P. H. Hayward, and E. O. Garton. 1993. Ecology of boreal owls in the northern Rocky Mountains, USA. Wildl. Mono. 59 pp.


Hayward, G. D., R. K. Steinhorst, and P. H. Hayward. 1992. Monitoring boreal owl populations with nest boxes: Sample size and cost. J. Wildl. Manage. 56:777-785.


Hayward, G. D. 1991. Using population biology to define old-growth forest. Wildl. Soc. Bull. 19:111-116.


Hayward, G. D., C. B. Kepler, and J. M. Scott. 1991. Point counts from clustered populations: Lessons from an experiment with Hawaiian Crows. Condor 93:676-682.


Hayward, G. D. 1989. Historical grizzly bear trends in Glacier National Park, Montana: A critique. Wildl. Soc. Bull. 17:195-197.


Hayward, P. H. and G. D. Hayward. 1989. Lone rangers of the Rockies. Natural History Magazine, November 1989:79-85.


Hayward, G. D. and R. E. Escano. 1989. Goshawk nest-site characteristics in western Montana and northern Idaho. Condor 91:476-479.


Hayward, G. D. and E. O. Garton. 1988. Resource partitioning among forest owls in the River of No Return Wilderness, Idaho. Oecologia 75:253-265.